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Page last updated
Sat 17 Feb 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea

family UNIONIDAE Rafinesque, 1820: cladomics
     5 subfamilies, 33 genera

Cladomics Summary

number of species = 1036
number of subtaxa (rank: subfamily) = 38

analyzed in 492 cladograms in 275 publications
maximum number of terminals in an analysis = 204, mean = 21.11
maximum number of subtaxa (subfamily) in an analysis = 4, mean = 1.91

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 470
maximum number of included terminals = 204, mean = 22.05
maximum number of subtaxa (subfamily) = 4, mean = 1.96
proportion of cladograms that support basic monophyly = 0.92

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 171
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (subfamily) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 204, mean = 30.84
maximum number of subtaxa (subfamily) = 4, mean = 2.92
proportion of cladograms that support rigorous monophyly = 0.82


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree


Hoeh et al., 2009 [Publ 2716], fig. 5

iT tT bi riS oT roT roS iD
24 45 39 4 22 2 1 0.0

*** Top Monophyly Cladogram ***

  +----------------------------- Coelatura_aegyptiaca **       
  |                                                         
  |                          +-- Margaritifera_margaritifera
  |     +--------------------+                              
  |     |                    +-- Cumberlandia_monodonta     
  |     |                                                   
  |     |                 +----- Velesunio_ambiguus         
  |     |        +--------+                                 
  |     |        |        |  +-- Velesunio_angasi           
  |     |        |        +--+                              
  |  +--+  +-----+           +-- Lortiella_rugata           
  |  |  |  |     |                                          
  |  |  |  |     |  +----------- Echyridella_menziesii      
  |  |  |  |     |  |                                       
--+  |  |  |     +--+        +-- Hyridella_depressa         
  |  |  |  |        |  +-----+                              
  |  |  |  |        |  |     +-- Hyridella_australis        
  |  |  |  |        +--+                                    
  |  |  +--+           |  +----- Castalia                   
  |  |     |           +--+                                 
  |  |     |              |  +-- Diplodon_chilensis         
  |  |     |              +--+                              
  |  |     |                 +-- Diplodon_gratus            
  |  |     |                                                
  |  |     |  +----------------- Pseudomulleria_dalyi       
  |  |     |  |                                             
  |  |     +--+  +-------------- Mulleria_rivoli            
  |  |        |  |                                          
  |  |        +--+  +----------- Etheria_elliptica          
  +--+           |  |                                       
     |           |  |     +----- Monocondylaea_minuana      
     |           +--+  +--+                                 
     |              |  |  |  +-- Anodontites_trigona        
     |              |  |  +--+                              
     |              |  |     +-- Anodontites_guanarensis    
     |              +--+                                    
     |                 |     +-- Mutela_rostrata            
     |                 |  +--+                              
     |                 |  |  +-- Mutela_dubia               
     |                 +--+                                 
     |                    |  +-- Chambardia_wahlbergi       
     |                    +--+                              
     |                       +-- Aspatharia_rugifera        
     |                                                      
     |                       +-- Pilsbryoconcha_exilis **      
     |  +--------------------+                              
     |  |                    +-- Pseudodon_vondembuschianus ** 
     |  |                                                   
     |  |  +-------------------- Lens_contradens **            
     +--+  |                                                
        |  |                 +-- Unio_pictorum **              
        |  |        +--------+                              
        |  |        |        +-- Unio_caffer **                
        +--+  +-----+                                       
           |  |     |  +-------- Lasmigona_costata **          
           |  |     +--+                                    
           |  |        |  +----- Alasmidonta_marginata **      
           |  |        +--+                                 
           |  |           |  +-- Strophitus_undulatus **       
           +--+           +--+                              
              |              +-- Pyganodon_grandis **          
              |                                             
              |  +-------------- Gonidea_angulata **           
              |  |                                          
              |  |        +----- Obliquaria_reflexa **         
              |  |  +-----+                                 
              +--+  |     |  +-- Quadrula_verrucosa **         
                 |  |     +--+                              
                 |  |        +-- Quadrula_quadrula **          
                 |  |                                       
                 +--+  +-------- Amblema_plicata **            
                    |  |                                    
                    |  |  +----- Eurynia_dilatata **           
                    |  +--+                                 
                    |  |  |  +-- Pleurobema_sintoxia **        
                    +--+  +--+                              
                       |     +-- Fusconaia_flava **            
                       |                                    
                       |  +----- Truncilla_truncata **         
                       |  |                                 
                       |  +----- Ptychobranchus_fasciolaris ** 
                       +--+                                 
                          |  +-- Ortmanniana_ligamentina **    
                          |  |                              
                          |  +-- Cambarunio_iris **            
                          +--+                              
                             +-- Ligumia_recta **              
                             |                              
                             +-- Epioblasma_triquetra **       

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 25 14 1 46.0

*** Top Sister-Group Cladogram ***

                 +----- Potomida_littoralis **        
  +--------------+                                 
  |              |  +-- Lampsilis_cardium **          
  |              +--+                              
  |                 +-- Unio_pictorum **              
  |                                                
--+              +----- Gibbosula_rochechouartii   
  |  +-----------+                                 
  |  |           |  +-- Gibbosula_crassa           
  |  |           +--+                              
  |  |              +-- Gibbosula_laosensis        
  |  |                                             
  +--+        +-------- Cumberlandia_monodonta     
     |  +-----+                                    
     |  |     |  +----- Pseudunio_marocanus        
     |  |     +--+                                 
     |  |        |  +-- Pseudunio_auricularius     
     |  |        +--+                              
     +--+           +-- Pseudunio_homsensis        
        |                                          
        |           +-- Margaritifera_dahurica     
        |  +--------+                              
        |  |        +-- Margaritifera_margaritifera
        +--+                                       
           |  +-------- Margaritifera_falcata      
           |  |                                    
           +--+     +-- Margaritifera_middendorffi 
              |  +--+                              
              |  |  +-- Margaritifera_laevis       
              +--+                                 
                 |  +-- Margaritifera_hembeli      
                 +--+                              
                    +-- Margaritifera_marrianae    

Inoue et al., 2018 [Publ 3858], fig. 1, 3a-c

iT tT bi riS oT roT roS iD
204 204 65 3 1 1 1 210.0
  +-------------------------------------------------------- Cumberlandia_monodonta     
  |                                                                                    
  |                                                     +-- Alasmidonta_marginata **      
  |                                                     |                              
  |                                                     +-- Prolasmidonta_heterodon **    
  |                                                     |                              
  |                                                     +-- Alasmidonta_undulata **       
  |                                                     |                              
  |                                                     +-- Beringiana_beringiana **      
  |                                                     |                              
  |                                                     +-- Anodonta_kennerlyi **         
  |                                                     |                              
  |                                                     +-- Anodonta_nuttalliana **       
  |                                                     |                              
  |                                                     +-- Anodonta_oregonensis **       
  |                                                     |                              
  |                                                     +-- Arcidens_confragosus **       
  |                                                     |                              
  |                                                     +-- Arcidens_wheeleri **          
  |                                                     |                              
--+                                                     +-- Lasmigona_complanata **       
  |                                                     |                              
  |  +--------------------------------------------------+-- Platynaias_compressa **       
  |  |                                                  |                              
  |  |                                                  +-- Lasmigona_costata **          
  |  |                                                  |                              
  |  |                                                  +-- Platynaias_decorata **        
  |  |                                                  |                              
  |  |                                                  +-- Platynaias_subviridis **      
  |  |                                                  |                              
  |  |                                                  +-- Pyganodon_grandis **          
  |  |                                                  |                              
  |  |                                                  +-- Pyganodon_lacustris **        
  |  |                                                  |                              
  |  |                                                  +-- Pseudodontoideus_subvexus **  
  |  |                                                  |                              
  |  |                                                  +-- Strophitus_undulatus **       
  |  |                                                  |                              
  |  |                                                  +-- Utterbackia_imbecillis **     
  |  |                                                  |                              
  +--+                                                  +-- Utterbackia_peninsularis[1] **
     |                                                  |                              
     |                                                  +-- Utterbackia_peninsularis[2] **
     |                                                                                 
     |  +-------------------------------------------------- Gonidea_angulata **           
     |  |                                                                              
     |  |                                               +-- Reginaia_ebenus **            
     |  |  +--------------------------------------------+                              
     |  |  |                                            +-- Reginaia_rotulata **          
     |  |  |                                                                           
     |  |  |                                            +-- Amblema_plicata **            
     |  |  +--------------------------------------------+                              
     |  |  |                                            +-- Amblema_elliottii **          
     |  |  |                                                                           
     |  |  |                                         +----- Popenaias_popeii **           
     |  |  |                                         |                                 
     |  |  |                                         |  +-- Dromus_dromas **              
     |  |  |                                         |  |                              
     +--+  |                                         |  +-- Epioblasma_rangiana **        
        |  |                                         |  |                              
        |  +-----------------------------------------+  +-- Glebula_rotundata **          
        |  |                                         |  |                              
        |  |                                         |  +-- Lampsilis_ovata **            
        |  |                                         |  |                              
        |  |                                         |  +-- Lampsilis_virescens **        
        |  |                                         |  |                              
        |  |                                         |  +-- Lemiox_rimosus **             
        |  |                                         +--+                              
        |  |                                            +-- Obliquaria_reflexa **         
        |  |                                            |                              
        |  |                                            +-- Obovaria_olivaria **          
        |  |                                            |                              
        +--+                                            +-- Obovaria_unicolor **          
           |                                            |                              
           |                                            +-- Plectomerus_dombeyanus **     
           |                                            |                              
           |                                            +-- Potamilus_alatus **           
           |                                            |                              
           |                                            +-- Potamilus_capax **            
           |                                                                           
           |                                            +-- Uniomerus_declivis **         
           |                                         +--+                              
           |                                         |  +-- Uniomerus_tetralasmus **      
           |                                         |                                 
           |                                         |  +-- Cyclonaias_tuberculata **     
           |  +--------------------------------------+  |                              
           |  |                                      |  +-- Pustulosa_pustulosa[1] **     
           |  |                                      |  |                              
           |  |                                      |  +-- Theliderma_metanevra **       
           |  |                                      +--+                              
           |  |                                         +-- Pustulosa_pustulosa[2] **     
           |  |                                         |                              
           |  |                                         +-- Quadrula_quadrula **          
           |  |                                         |                              
           |  |                                         +-- Quadrula_verrucosa **         
           |  |                                                                        
           |  |  +----------------------------------------- Elliptoideus_sloatianus **    
           +--+  |                                                                     
              |  |                                      +-- Elliptio_lanceolata **        
              |  |  +-----------------------------------+                              
              |  |  |                                   +-- Parvaspina_collina **         
              |  |  |                                                                  
              |  |  |  +----------------------------------- Plethobasus_cyphyus **        
              |  |  |  |                                                               
              |  |  |  +----------------------------------- Eurynia_dilatata **           
              |  |  |  |                                                               
              |  |  |  +----------------------------------- Hemistena_lata **             
              |  |  |  |                                                               
              |  |  |  |                                +-- Elliptio_complanata **        
              +--+  |  |                                |                              
                 |  |  |                                +-- Elliptio_hopetonensis **      
                 |  |  |                                |                              
                 |  |  |                                +-- Elliptio_producta[1] **       
                 |  |  |                                |                              
                 |  |  |                                +-- Elliptio_shepardiana **       
                 |  |  |                                |                              
                 |  |  |                                +-- Elliptio_angustata **         
                 |  |  |                                |                              
                 |  |  |                                +-- Elliptio_arca **              
                 |  |  |                                |                              
                 |  |  |                                +-- Elliptio_arctata **           
                 |  |  |                                |                              
                 +--+  |                                +-- Elliptio_jayensis **          
                    |  |                                |                              
                    |  |                                +-- Elliptio_crassidens **        
                    |  |                                |                              
                    |  +--------------------------------+-- Elliptio_dariensis **         
                    |  |                                |                              
                    |  |                                +-- Elliptio_fisheriana **        
                    |  |                                |                              
                    |  |                                +-- Elliptio_folliculata **       
                    |  |                                |                              
                    |  |                                +-- Elliptio_fumata **            
                    |  |                                |                              
                    |  |                                +-- Elliptio_icterina **          
                    |  |                                |                              
                    |  |                                +-- Elliptio_mcmichaeli **        
                    |  |                                |                              
                    |  |                                +-- Elliptio_producta[2] **       
                    |  |                                |                              
                    |  |                                +-- Elliptio_pullata **           
                    |  |                                |                              
                    |  |                                +-- Elliptio_purpurella **        
                    |  |                                |                              
                    |  |                                +-- Elliptio_congaraea **         
                    |  |                                                               
                    +--+                                +-- Pleuronaia_barnesiana **      
                       |                                |                              
                       |  +-----------------------------+-- Pleuronaia_dolabelloides **   
                       |  |                             |                              
                       |  |                             +-- Pleuronaia_gibber **          
                       |  |                                                            
                       |  |     +-------------------------- Pleurobema_plenum **          
                       |  |     |                                                      
                       |  |     |  +----------------------- Pleurobema_stabile **         
                       |  |     |  |                                                   
                       |  |     |  |                 +----- Pleurobema_pyriforme **       
                       |  |  +--+  |              +--+                                 
                       |  |  |  |  |              |  |  +-- Pleurobema_georgianum[2] **   
                       |  |  |  |  |              |  +--+                              
                       |  |  |  |  |  +-----------+     +-- Pleurobema_athearni **        
                       |  |  |  |  |  |           |                                    
                       |  |  |  +--+  |           |  +----- Pleurobema_decisum **         
                       |  |  |     |  |           |  |                                 
                       |  |  |     |  |           +--+  +-- Pleurobema_georgianum[1] **   
                       |  |  |     |  |              |  |                              
                       |  |  |     |  |              +--+-- Pleurobema_hanleyianum[1] **  
                       |  |  |     |  |                 |                              
                       |  |  |     |  |                 +-- Pleurobema_hanleyianum[2] **  
                       |  |  |     |  |                                                
                       |  |  |     +--+     +-------------- Pleurobema_beadleianum[2] **  
                       |  |  |        |     |                                          
                       |  |  |        |  +--+  +----------- Pleurobema_strodeanum[2] **   
                       |  |  |        |  |  |  |                                       
                       |  |  |        |  |  +--+----------- Pleurobema_rubellum **        
                       |  |  |        |  |     |                                       
                       |  |  |        |  |     |  +-------- Pleurobema_beadleianum[1] **  
                       |  |  |        |  |     +--+                                    
                       |  |  |        |  |        |  +----- Pleurobema_taitianum **       
                       |  |  |        |  |        +--+                                 
                       |  |  |        |  |           |  +-- Pleurobema_perovatum **       
                       |  |  |        +--+           +--+                              
                       |  |  |           |              +-- Pleurobema_strodeanum[1] **   
                       |  |  |           |                                             
                       |  |  |           |           +----- Pleurobema_oviforme[6] **     
                       |  |  |           |           |                                 
                       |  |  |           |        +--+----- Pleurobema_clava[3] **        
                       |  |  |           |        |  |                                 
                       |  |  |           |        |  |  +-- Pleurobema_oviforme[5] **     
                       |  |  |           |        |  +--+                              
                       |  |  |           +--------+     +-- Pleurobema_clava[2] **        
                       +--+  |                    |                                    
                          |  |                    |  +----- Pleurobema_oviforme[2] **     
                          |  |                    |  |                                 
                          |  |                    |  +----- Pleurobema_oviforme[3] **     
                          |  |                    +--+                                 
                          |  |                       +----- Pleurobema_oviforme[4] **     
                          |  |                       |                                 
                          |  |                       |  +-- Pleurobema_clava[1] **        
                          |  |                       +--+                              
                          |  |                          +-- Pleurobema_oviforme[1] **     
                          |  |                                                         
                          |  |  +-------------------------- Pleurobema_cordatum **        
                          |  |  |                                                      
                          |  |  |                    +----- Fusconaia_subrotunda **       
                          |  |  |  +-----------------+                                 
                          |  +--+  |                 |  +-- Fusconaia_cor **              
                          |  |  |  |                 +--+                              
                          |  |  |  |                    +-- Fusconaia_cuneolus **         
                          |  |  |  |                                                   
                          |  |  |  |  +-------------------- Fusconaia_ozarkensis **       
                          |  |  +--+  |                                                
                          |  |     |  |                 +-- Fusconaia_escambia **         
                          |  |     |  |  +--------------+                              
                          |  |     |  |  |              +-- Fusconaia_burkei **           
                          |  |     |  |  |                                             
                          |  |     |  |  |           +----- Fusconaia_flava[27] **        
                          |  |     |  |  |           |                                 
                          |  |     +--+  |           |  +-- Fusconaia_iheringi **         
                          |  |        |  |           +--+                              
                          |  |        |  |           |  +-- Fusconaia_mitchelli **        
                          |  |        |  +-----------+                                 
                          |  |        |  |           |  +-- Fusconaia_askewi[1] **        
                          |  |        |  |           |  |                              
                          |  |        |  |           |  +-- Fusconaia_askewi[2] **        
                          |  |        |  |           |  |                              
                          |  |        +--+           |  +-- Fusconaia_askewi[3] **        
                          |  |           |           +--+                              
                          |  |           |              +-- Fusconaia_askewi[4] **        
                          |  |           |              |                              
                          |  |           |              +-- Fusconaia_askewi[5] **        
                          |  |           |              |                              
                          |  |           |              +-- Fusconaia_chunii **           
                          |  |           |                                             
                          +--+           |  +-------------- Fusconaia_masoni **           
                             |           |  |                                          
                             |           |  |  +----------- Fusconaia_flava[26] **        
                             |           |  |  |                                       
                             |           |  |  |        +-- Fusconaia_cerina[8] **        
                             |           +--+  |  +-----+                              
                             |              |  |  |     +-- Fusconaia_flava[25] **        
                             |              |  |  |                                    
                             |              |  |  |  +----- Fusconaia_flava[5] **         
                             |              |  |  |  |                                 
                             |              |  |  |  +----- Fusconaia_flava[6] **         
                             |              +--+  |  |                                 
                             |                 |  |  +----- Fusconaia_flava[7] **         
                             |                 |  |  |                                 
                             |                 |  |  +----- Fusconaia_flava[8] **         
                             |                 |  |  |                                 
                             |                 |  |  +----- Fusconaia_cerina[4] **        
                             |                 |  |  |                                 
                             |                 |  |  +----- Fusconaia_flava[9] **         
                             |                 |  |  |                                 
                             |                 +--+  +----- Fusconaia_flava[10] **        
                             |                    |  |                                 
                             |                    |  +----- Fusconaia_flava[11] **        
                             |                    |  |                                 
                             |                    |  +----- Fusconaia_flava[12] **        
                             |                    |  |                                 
                             |                    |  +----- Fusconaia_flava[13] **        
                             |                    |  |                                 
                             |                    |  +----- Fusconaia_flava[14] **        
                             |                    |  |                                 
                             |                    |  +----- Fusconaia_flava[15] **        
                             |                    |  |                                 
                             |                    |  +----- Fusconaia_flava[16] **        
                             |                    |  |                                 
                             |                    |  +----- Fusconaia_flava[17] **        
                             |                    +--+                                 
                             |                       +----- Fusconaia_flava[18] **        
                             |                       |                                 
                             |                       +----- Fusconaia_flava[19] **        
                             |                       |                                 
                             |                       +----- Fusconaia_flava[20] **        
                             |                       |                                 
                             |                       +----- Fusconaia_flava[21] **        
                             |                       |                                 
                             |                       +----- Fusconaia_flava[22] **        
                             |                       |                                 
                             |                       +----- Fusconaia_flava[23] **        
                             |                       |                                 
                             |                       +----- Fusconaia_flava[24] **        
                             |                       |                                 
                             |                       +----- Fusconaia_cerina[5] **        
                             |                       |                                 
                             |                       +----- Fusconaia_cerina[6] **        
                             |                       |                                 
                             |                       +----- Fusconaia_cerina[7] **        
                             |                       |                                 
                             |                       |  +-- Fusconaia_flava[1] **         
                             |                       |  |                              
                             |                       |  +-- Fusconaia_flava[2] **         
                             |                       |  |                              
                             |                       |  +-- Fusconaia_flava[3] **         
                             |                       |  |                              
                             |                       +--+-- Fusconaia_flava[4] **         
                             |                          |                              
                             |                          +-- Fusconaia_cerina[1] **        
                             |                          |                              
                             |                          +-- Fusconaia_cerina[2] **        
                             |                          |                              
                             |                          +-- Fusconaia_cerina[3] **        
                             |                                                         
                             |           +----------------- Pleurobema_riddellii **       
                             |           |                                             
                             |           |  +-------------- Pleurobema_sintoxia[21] **    
                             |           |  |                                          
                             |           |  |  +----------- Pleurobema_sintoxia[19] **    
                             |           |  |  |                                       
                             |           |  |  +----------- Pleurobema_sintoxia[20] **    
                             |           |  |  |                                       
                             +-----------+  |  |     +----- Pleurobema_sintoxia[18] **    
                                         |  |  |     |                                 
                                         |  |  |     +----- Pleurobema_rubrum[19] **      
                                         |  |  |     |                                 
                                         |  |  |     +----- Pleurobema_rubrum[20] **      
                                         |  |  +-----+                                 
                                         |  |  |     +----- Pleurobema_rubrum[21] **      
                                         |  |  |     |                                 
                                         |  |  |     |  +-- Pleurobema_rubrum[17] **      
                                         +--+  |     |  |                              
                                            |  |     |  +-- Pleurobema_sintoxia[16] **    
                                            |  |     +--+                              
                                            |  |        +-- Pleurobema_sintoxia[17] **    
                                            |  |        |                              
                                            |  |        +-- Pleurobema_rubrum[18] **      
                                            |  |                                       
                                            |  |  +-------- Pleurobema[4] **              
                                            |  |  |                                    
                                            |  |  +-------- Pleurobema[5] **              
                                            |  |  |                                    
                                            |  |  +-------- Pleurobema_sintoxia[1] **     
                                            |  |  |                                    
                                            |  |  +-------- Pleurobema[8] **              
                                            |  |  |                                    
                                            |  |  +-------- Pleurobema[9] **              
                                            +--+  |                                    
                                               |  +-------- Pleurobema[10] **             
                                               |  |                                    
                                               |  +-------- Pleurobema_sintoxia[3] **     
                                               |  |                                    
                                               |  +-------- Pleurobema_sintoxia[4] **     
                                               |  |                                    
                                               |  +-------- Pleurobema_sintoxia[5] **     
                                               |  |                                    
                                               |  +-------- Pleurobema_sintoxia[6] **     
                                               |  |                                    
                                               |  +-------- Pleurobema_sintoxia[7] **     
                                               |  |                                    
                                               |  +-------- Pleurobema_sintoxia[8] **     
                                               |  |                                    
                                               |  +-------- Pleurobema_sintoxia[9] **     
                                               |  |                                    
                                               |  +-------- Pleurobema_sintoxia[10] **    
                                               |  |                                    
                                               |  +-------- Pleurobema_sintoxia[11] **    
                                               |  |                                    
                                               |  +-------- Pleurobema_sintoxia[12] **    
                                               |  |                                    
                                               |  +-------- Pleurobema_sintoxia[13] **    
                                               |  |                                    
                                               |  +-------- Pleurobema_sintoxia[14] **    
                                               |  |                                    
                                               |  +-------- Pleurobema_sintoxia[15] **    
                                               |  |                                    
                                               +--+-------- Pleurobema_rubrum[4] **       
                                                  |                                    
                                                  +-------- Pleurobema_rubrum[5] **       
                                                  |                                    
                                                  +-------- Pleurobema_rubrum[6] **       
                                                  |                                    
                                                  +-------- Pleurobema_rubrum[7] **       
                                                  |                                    
                                                  +-------- Pleurobema_rubrum[8] **       
                                                  |                                    
                                                  +-------- Pleurobema_rubrum[9] **       
                                                  |                                    
                                                  +-------- Pleurobema_rubrum[10] **      
                                                  |                                    
                                                  +-------- Pleurobema_rubrum[11] **      
                                                  |                                    
                                                  +-------- Pleurobema_rubrum[12] **      
                                                  |                                    
                                                  +-------- Pleurobema_rubrum[13] **      
                                                  |                                    
                                                  +-------- Pleurobema_rubrum[14] **      
                                                  |                                    
                                                  +-------- Pleurobema_rubrum[15] **      
                                                  |                                    
                                                  +-------- Pleurobema_rubrum[16] **      
                                                  |                                    
                                                  |     +-- Pleurobema[6] **              
                                                  +-----+                              
                                                  |     +-- Pleurobema_rubrum[1] **       
                                                  |                                    
                                                  |     +-- Pleurobema_sintoxia[2] **     
                                                  +-----+                              
                                                  |     +-- Pleurobema_rubrum[2] **       
                                                  |                                    
                                                  |     +-- Pleurobema[7] **              
                                                  +-----+                              
                                                  |     +-- Pleurobema_rubrum[3] **       
                                                  |                                    
                                                  |  +----- Pleurobema[3] **              
                                                  +--+                                 
                                                     |  +-- Pleurobema[1] **              
                                                     +--+                              
                                                        +-- Pleurobema[2] **              

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
165 165 142 4 39 13 1 180.0
                             +----------------- Gibbosula_rochechouartii   
                             |                                             
                             |  +-------------- Gibbosula_laosensis        
  +--------------------------+  |                                          
  |                          |  |  +----------- Cumberlandia_monodonta     
  |                          |  |  |                                       
  |                          +--+  |     +----- Pseudunio_marocanus        
  |                             |  +-----+                                 
  |                             |  |     |  +-- Pseudunio_auricularius     
  |                             |  |     +--+                              
  |                             +--+        +-- Pseudunio_homsensis        
  |                                |                                       
  |                                |        +-- Margaritifera_dahurica     
  |                                |  +-----+                              
  |                                |  |     +-- Margaritifera_margaritifera
  |                                |  |                                    
  |                                +--+  +----- Margaritifera_laevis       
  |                                   |  |                                 
  |                                   |  |  +-- Margaritifera_falcata      
  |                                   +--+--+                              
  |                                      |  +-- Margaritifera_middendorffi 
  |                                      |                                 
  |                                      |  +-- Margaritifera_hembeli      
  |                                      +--+                              
  |                                         +-- Margaritifera_marrianae    
  |                                                                        
  |                                      +----- Trapezidens_exolescens **     
  |                    +-----------------+                                 
  |                    |                 |  +-- Trapezidens_dolichorhynchus **
  |                    |                 +--+                              
  |                 +--+                    +-- Trapezidens_angustior **      
  |                 |  |                                                   
  |                 |  |  +-------------------- Lamellidens_corrianus **      
  |                 |  |  |                                                
  |                 |  +--+  +----------------- Lamellidens_savadiensis **    
  |                 |     |  |                                             
  |                 |     +--+              +-- Lamellidens_lamellatus **     
  |                 |        |  +-----------+                              
  |                 |        |  |           +-- Lamellidens_jenkinsianus **   
  |                 |        +--+                                          
  |                 |           |  +----------- Lamellidens_generosus **      
  |                 |           +--+                                       
  |                 |              |  +-------- Lamellidens_mainwaringi[2] ** 
  |                 |              +--+                                    
  |  +--------------+                 |  +----- Lamellidens_mainwaringi[1] ** 
  |  |              |                 +--+                                 
  |  |              |                    |  +-- Lamellidens_ferrugineus **    
  |  |              |                    +--+                              
  |  |              |                       +-- Lamellidens_marginalis **     
  |  |              |                                                      
  |  |              |                    +----- Parreysia_keralaensis **      
  |  |              |  +-----------------+                                 
  |  |              |  |                 |  +-- Parreysia_nagpoorensis **     
  |  |              |  |                 +--+                              
  |  |              |  |                    +-- Parreysia_plagiosoma **       
--+  |              |  |                                                   
  |  |              |  |                    +-- Leoparreysia_tavoyensis **    
  |  |              |  |                    |                              
  |  |              +--+  +-----------------+-- Leoparreysia_canefrii **      
  |  |                 |  |                 |                              
  |  |                 |  |                 +-- Leoparreysia_olivacea **      
  |  |                 |  |                                                
  |  |                 |  |           +-------- Coelatura_aegyptiaca[2] **    
  |  |                 |  |           |                                    
  |  |                 |  |  +--------+  +----- Coelatura_aegyptiaca[1] **    
  |  |                 +--+  |        |  |                                 
  |  |                    |  |        |  +----- Nitia_teretiuscula **         
  |  |                    |  |        +--+                                 
  |  |                    |  |           |  +-- Coelatura_gabonensis **       
  |  |                    |  |           |  |                              
  |  |                    |  |           +--+-- Coelatura_kunenensis **       
  |  |                    |  |              |                              
  |  |                    +--+              +-- Prisodontopsis_mweruensis **  
  |  |                       |                                             
  |  |                       |        +-------- Indonaia_shurtleffiana **     
  |  |                       |        |                                    
  |  |                       |  +-----+  +----- Indonaia_bonneaudii **        
  |  |                       |  |     |  |                                 
  |  |                       |  |     +--+----- Indonaia_rugosa **            
  |  |                       |  |        |                                 
  |  |                       |  |        |  +-- Indonaia_andersoniana **      
  |  |                       +--+        +--+                              
  |  |                          |           +-- Indonaia_caerulea **          
  |  |                          |                                          
  |  |                          |  +----------- Indochinella_pugio **         
  |  |                          |  |                                       
  |  |                          |  |     +----- Scabies_humilis **            
  |  |                          +--+  +--+                                 
  |  |                             |  |  |  +-- Scabies_phaselus **           
  |  |                             |  |  +--+                              
  |  |                             +--+     +-- Scabies_mandarinus **         
  |  |                                |                                    
  |  |                                |  +----- Radiatula_myitkyinae **       
  |  |                                +--+                                 
  |  |                                   |  +-- Radiatula_chaudhurii **       
  |  |                                   +--+                              
  |  |                                      +-- Radiatula_mouhoti **          
  |  |                                                                     
  |  |        +-------------------------------- Acuticosta_chinensis **       
  |  |        |                                                            
  |  |        |  +----------------------------- Lepidodesma_languilati **     
  |  |        |  |                                                         
  |  |        |  |                    +-------- Aculamprotula_tortuosa **     
  |  |        |  |                    |                                    
  |  |        |  |        +-----------+  +----- Aculamprotula_coreana **      
  |  |        |  |        |           |  |                                 
  |  |        |  |        |           +--+----- Aculamprotula_scripta **      
  |  |        |  |        |              |                                 
  +--+        |  |        |              |  +-- Aculamprotula_fibrosa **      
     |        |  |        |              +--+                              
     |        |  |        |                 +-- Aculamprotula_tientsinensis **
     |  +-----+  +--------+                                                
     |  |     |  |        |           +-------- Unio_tumidus **               
     |  |     |  |        |  +--------+                                    
     |  |     |  |        |  |        |  +----- Unio_crassus **               
     |  |     |  |        |  |        +--+                                 
     |  |     |  |        |  |           |  +-- Unio_delphinus **             
     |  |     |  |        |  |           +--+                              
     |  |     |  |        +--+              +-- Unio_pictorum **              
     |  |     |  |           |                                             
     |  |     |  |           |              +-- Schistodesmus_lampreyanus **  
     |  |     |  |           |  +-----------+                              
     |  |     |  |           |  |           +-- Schistodesmus_spinosus **     
     |  |     |  |           |  |                                          
     |  |     |  |           +--+        +----- Nodularia_douglasiae[1] **    
     |  |     +--+              |  +-----+                                 
     |  |        |              |  |     |  +-- Nodularia_douglasiae[2] **    
     |  |        |              |  |     +--+                              
     |  |        |              +--+        +-- Nodularia_nuxpersicae **      
     |  |        |                 |                                       
     |  |        |                 |  +-------- Tchangsinaia_piscicula **     
     |  |        |                 +--+                                    
     |  |        |                    |  +----- Cuneopsis_rufescens **        
     |  |        |                    +--+                                 
     |  |        |                       |  +-- Cuneopsis_heudei **           
     |  |        |                       +--+                              
     |  |        |                          +-- Cuneopsis_celtiformis **      
     |  |        |                                                         
     |  |        |                    +-------- Lanceolaria_gladiola **       
     |  |        |                    |                                    
     |  |        |  +-----------------+  +----- Lanceolaria_grayii[1] **      
     |  |        |  |                 |  |                                 
     |  |        |  |                 +--+----- Lanceolaria_grayii[2] **      
     |  |        |  |                    |                                 
     |  |        |  |                    |  +-- Lanceolaria_lanceolata **     
     |  |        |  |                    +--+                              
     |  |        |  |                       +-- Lanceolaria_triformis **      
     |  |        |  |                                                      
     |  |        |  |     +-------------------- Anemina_arcaeformis **        
     |  |        |  |     |                                                
     |  |        |  |  +--+  +----------------- Cristaria_plicata **          
     |  |        +--+  |  |  |                                             
     |  |           |  |  +--+  +-------------- Pletholophus_tenuis **        
     |  |           |  |     |  |                                          
     |  |           |  |     +--+           +-- Sinanodonta_angula **         
     |  |           |  |        |  +--------+                              
     |  |           |  |        |  |        +-- Sinanodonta_lucida **         
     |  |           |  |        +--+                                       
     |  |           |  |           |  +-------- Sinanodonta_woodiana[2] **    
     |  |           |  |           +--+                                    
     |  |           |  |              |  +----- Sinanodonta_jourdyi **        
     |  |           |  |              +--+                                 
     |  |           +--+                 |  +-- Sinanodonta_elliptica **      
     |  |              |                 +--+                              
     |  |              |                    +-- Sinanodonta_woodiana[1] **    
     |  |              |                                                   
     |  |              |              +-------- Alasmidonta_marginata **      
     |  |              |              |                                    
     |  |              |              +-------- Lasmigona_costata **          
     |  |              |              |                                    
     |  |              |              +-------- Pseudanodonta_complanata **   
     |  |              |              |                                    
     |  |              |              +-------- Simpsonaias_ambigua **        
     +--+              |              |                                    
        |              +--------------+-------- Strophitus_undulatus **       
        |                             |                                    
        |                             |  +----- Anodonta_anatina **           
        |                             +--+                                 
        |                             |  |  +-- Anodonta_cygnea **            
        |                             |  +--+                              
        |                             |     +-- Anodonta_nuttalliana **       
        |                             |                                    
        |                             |  +----- Pyganodon_grandis **          
        |                             +--+                                 
        |                                |  +-- Utterbackia_imbecillis **     
        |                                +--+                              
        |                                   +-- Utterbackia_peninsularis **   
        |                                                                  
        |                          +----------- Hyriopsis_desowitzi **        
        |                          |                                       
        |                          +----------- Rectidens_sumatrensis **      
        |                          |                                       
        |                          |        +-- Hyriopsis_bialata **          
        |                          |        |                              
        |                          |        +-- Hyriopsis_myersiana **        
        |                    +-----+--------+                              
        |                    |     |        +-- Hyriopsis_panhai **           
        |                    |     |        |                              
        |                    |     |        +-- Hyriopsis_khoratensis **      
        |                    |     |                                       
        |                    |     |        +-- Ensidens_ingallsianus **      
        |                    |     |  +-----+                              
        |                    |     |  |     +-- Ensidens_telus **             
        |                    |     +--+                                    
        |                    |        |  +----- Ensidens_sagittarius **       
        |  +-----------------+        +--+                                 
        |  |                 |           |  +-- Ensidens_spiculus **          
        |  |                 |           +--+                              
        |  |                 |              +-- Ensidens_jaculus **           
        |  |                 |                                             
        |  |                 |           +----- Physunio **                   
        |  |                 |  +--------+                                 
        |  |                 |  |        |  +-- Physunio_modelli **           
        |  |                 |  |        +--+                              
        |  |                 |  |           +-- Physunio_superbus **          
        |  |                 |  |                                          
        |  |                 +--+     +-------- Yaukthwa_inlenensis **        
        |  |                    |  +--+                                    
        |  |                    |  |  |  +----- Yaukthwa_panhai **            
        |  |                    |  |  +--+                                 
        |  |                    |  |     |  +-- Yaukthwa_nesemanni **         
        |  |                    |  |     +--+                              
        |  |                    +--+        +-- Yaukthwa_rectangularis **     
        |  |                       |                                       
        |  |                       |        +-- Lens_contradens **            
        |  |                       |  +-----+                              
        |  |                       |  |     +-- Lens_micropterus **           
        |  |                       +--+                                    
        |  |                          |  +----- Lens_eximius **               
        |  |                          +--+                                 
        |  |                             |  +-- Lens_comptus **               
        |  |                             +--+                              
        |  |                                +-- Lens_novoselovi **            
        |  |                                                               
        |  |                                +-- Megalonaias_nervosa **        
        |  |                             +--+                              
        +--+                             |  +-- Uniomerus_carolinianus **     
           |     +-----------------------+                                 
           |     |                       |  +-- Quadrula_quadrula **          
           |     |                       +--+                              
           |     |                          +-- Quadrula_verrucosa **         
           |     |                                                         
           |     |                    +-------- Elliptoideus_sloatianus **    
           |  +--+                    |                                    
           |  |  |  +-----------------+     +-- Elliptio_complanata **        
           |  |  |  |                 |  +--+                              
           |  |  |  |                 |  |  +-- Eurynia_dilatata **           
           |  |  |  |                 +--+                                 
           |  |  |  |                    |  +-- Fusconaia_flava **            
           |  |  +--+                    +--+                              
           |  |     |                       +-- Pleurobema_sintoxia **        
           |  |     |                                                      
           |  |     |  +----------------------- Amblema_plicata **            
           |  |     |  |                                                   
           |  |     |  |                 +----- Toxolasma_parvum **           
           |  |     +--+  +--------------+                                 
           |  |        |  |              |  +-- Obliquaria_reflexa **         
           |  |        |  |              +--+                              
           |  |        |  |                 +-- Plectomerus_dombeyanus **     
           |  |        +--+                                                
           |  |           |  +----------------- Potamilus_leptodon **         
           |  |           |  |                                             
           |  |           |  |  +-------------- Potamilus_alatus **           
           |  |           +--+  |                                          
           |  |              |  |  +----------- Venustaconcha_ellipsiformis **
           |  |              |  |  |                                       
           |  |              +--+  +----------- Leaunio_lienosus **           
           |  |                 |  |                                       
           +--+                 |  |        +-- Ellipsaria_lineolata **       
              |                 |  +--------+                              
              |                 +--+        +-- Truncilla_truncata **         
              |                    |                                       
              |                    |        +-- Medionidus_conradicus **      
              |                    +--------+                              
              |                    |        +-- Ptychobranchus_subtentus **   
              |                    |                                       
              |                    |  +-------- Cambarunio_iris **            
              |                    +--+                                    
              |                       |  +----- Ortmanniana_ligamentina **    
              |                       +--+                                 
              |                          |  +-- Lampsilis_cardium **          
              |                          +--+                              
              |                             +-- Lampsilis_ornata **           
              |                                                            
              |                          +----- Chamberlainia_somsakpanhai ** 
              |  +-----------------------+                                 
              |  |                       |  +-- Sinohyriopsis_cumingii **     
              |  |                       +--+                              
              |  |                          +-- Sinohyriopsis_schlegelii **   
              |  |                                                         
              |  |                          +-- Potomida_littoralis **        
              |  |                    +-----+                              
              |  |                    |     +-- Pronodularia_japanensis **    
              |  |  +-----------------+                                    
              +--+  |                 |  +----- Lamprotula_leaii **           
                 |  |                 +--+                                 
                 |  |                    |  +-- Lamprotula_caveata **         
                 |  |                    +--+                              
                 |  |                       +-- Lamprotula_cornuumlunae **    
                 |  |                                                      
                 |  |                 +-------- Sinosolenaia_carinata **      
                 |  |              +--+                                    
                 |  |              |  |  +----- Sinosolenaia_recognita **     
                 |  |              |  +--+                                 
                 +--+              |     |  +-- Sinosolenaia_oleivora **      
                    |              |     +--+                              
                    |  +-----------+        +-- Sinosolenaia_iridinea **      
                    |  |           |                                       
                    |  |           |     +----- Gonidea_angulata **           
                    |  |           |  +--+                                 
                    |  |           |  |  |  +-- Leguminaia_wheatleyi **       
                    |  |           |  |  +--+                              
                    |  |           +--+     +-- Microcondylaea_bonellii **    
                    |  |              |                                    
                    |  |              |     +-- Ptychorhynchus_pfisteri **    
                    |  |              |     |                              
                    +--+              +-----+-- Parvasolenaia_rivularis **    
                       |                    |                              
                       |                    +-- Koreosolenaia_sitgyensis **   
                       |                                                   
                       |                 +----- Indopseudodon_salwenianus **  
                       |  +--------------+                                 
                       |  |              |  +-- Indopseudodon_indawgyiensis **
                       |  |              +--+                              
                       |  |                 +-- Indopseudodon_bogani **       
                       |  |                                                
                       |  |           +-------- Pilsbryoconcha_exilis[3] **   
                       +--+  +--------+                                    
                          |  |        |  +----- Pilsbryoconcha_exilis[2] **   
                          |  |        +--+                                 
                          |  |           |  +-- Pilsbryoconcha_exilis[1] **   
                          |  |           +--+                              
                          |  |              +-- Pilsbryoconcha_carinifera **  
                          |  |                                             
                          +--+     +----------- Nyeinchanconcha_nyeinchani ** 
                             |  +--+                                       
                             |  |  |  +-------- Thaiconcha_callifera **       
                             |  |  +--+                                    
                             |  |     |  +----- Bineurus_loeiensis **         
                             |  |     +--+                                 
                             |  |        |  +-- Bineurus_mouhotii **          
                             +--+        +--+                              
                                |           +-- Bineurus_exilis **            
                                |                                          
                                |        +----- Pseudodon_vagulus **          
                                |     +--+                                 
                                |     |  |  +-- Pseudodon_mekongi **          
                                |     |  +--+                              
                                +-----+     +-- Pseudodon_vondembuschianus ** 
                                      |                                    
                                      |     +-- Sundadontina_cumingii **      
                                      |     |                              
                                      +-----+-- Sundadontina_tumida[1] **     
                                            |                              
                                            +-- Sundadontina_tumida[2] **     

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
163 163 97 4 39 13 1 177.0
           +-------------------- Gibbosula_rochechouartii   
           |                                                
  +--------+  +----------------- Gibbosula_laosensis        
  |        |  |                                             
  |        +--+  +-------------- Cumberlandia_monodonta     
  |           |  |                                          
  |           |  |        +----- Pseudunio_marocanus        
  |           +--+  +-----+                                 
  |              |  |     |  +-- Pseudunio_auricularius     
  |              |  |     +--+                              
  |              |  |        +-- Pseudunio_homsensis        
  |              +--+                                       
  |                 |  +-------- Margaritifera_falcata      
  |                 |  |                                    
  |                 |  |     +-- Margaritifera_dahurica     
  |                 |  +-----+                              
  |                 +--+     +-- Margaritifera_margaritifera
  |                    |                                    
  |                    |  +----- Margaritifera_laevis       
  |                    |  |                                 
  |                    +--+----- Margaritifera_middendorffi 
  |                       |                                 
  |                       |  +-- Margaritifera_hembeli      
  |                       +--+                              
  |                          +-- Margaritifera_marrianae    
  |                                                         
  |                          +-- Parreysia_keralaensis **      
  |                          |                              
  |     +--------------------+-- Parreysia_nagpoorensis **     
  |     |                    |                              
  |     |                    +-- Parreysia_plagiosoma **       
  |     |                                                   
  |     |                 +----- Trapezidens_exolescens **     
  |     |        +--------+                                 
  |     |        |        |  +-- Trapezidens_angustior **      
  |     |        |        +--+                              
  |     +--------+           +-- Trapezidens_dolichorhynchus **
  |     |        |                                          
  |     |        |  +----------- Lamellidens_corrianus **      
  |     |        |  |                                       
  |     |        +--+  +-------- Lamellidens_savadiensis **    
  |     |           |  |                                    
  |     |           |  |  +----- Lamellidens_marginalis **     
  |     |           +--+  |                                 
  |     |              |  +----- Lamellidens_ferrugineus **    
  |     |              |  |                                 
  |  +--+              +--+  +-- Lamellidens_mainwaringi[1] ** 
  |  |  |                 |  |                              
  |  |  |                 |  +-- Lamellidens_mainwaringi[2] ** 
  |  |  |                 |  |                              
  |  |  |                 +--+-- Lamellidens_lamellatus **     
  |  |  |                    |                              
  |  |  |                    +-- Lamellidens_generosus **      
  |  |  |                    |                              
  |  |  |                    +-- Lamellidens_jenkinsianus **   
  |  |  |                                                   
  |  |  |                 +----- Leoparreysia_tavoyensis **    
  |  |  |  +--------------+                                 
  |  |  |  |              |  +-- Leoparreysia_canefrii **      
  |  |  |  |              +--+                              
  |  |  |  |                 +-- Leoparreysia_olivacea **      
  |  |  |  |                                                
--+  |  |  |           +-------- Coelatura_aegyptiaca[2] **    
  |  |  |  |           |                                    
  |  |  |  |           +-------- Nitia_teretiuscula **         
  |  |  +--+  +--------+                                    
  |  |     |  |        |  +----- Coelatura_aegyptiaca[1] **    
  |  |     |  |        |  |                                 
  |  |     |  |        +--+  +-- Coelatura_gabonensis **       
  |  |     |  |           |  |                              
  |  |     |  |           +--+-- Coelatura_kunenensis **       
  |  |     |  |              |                              
  |  |     |  |              +-- Prisodontopsis_mweruensis **  
  |  |     +--+                                             
  |  |        |           +----- Indonaia_shurtleffiana **     
  |  |        |           |                                 
  |  |        |           |  +-- Indonaia_bonneaudii **        
  |  |        |  +--------+--+                              
  |  |        |  |        |  +-- Indonaia_rugosa **            
  |  |        |  |        |                                 
  |  |        |  |        |  +-- Indonaia_andersoniana **      
  |  |        |  |        +--+                              
  |  |        +--+           +-- Indonaia_caerulea **          
  |  |           |                                          
  |  |           |  +----------- Indochinella_pugio **         
  |  |           |  |                                       
  |  |           |  |     +----- Scabies_humilis **            
  |  |           +--+  +--+                                 
  |  |              |  |  |  +-- Scabies_phaselus **           
  |  |              |  |  +--+                              
  |  |              +--+     +-- Scabies_mandarinus **         
  |  |                 |                                    
  |  |                 |  +----- Radiatula_mouhoti **          
  |  |                 +--+                                 
  |  |                    |  +-- Radiatula_myitkyinae **       
  |  |                    +--+                              
  |  |                       +-- Radiatula_chaudhurii **       
  |  |                                                      
  |  |     +-------------------- Chamberlainia_somsakpanhai ** 
  |  |     |                                                
  |  |     +-------------------- Megalonaias_nervosa **        
  |  |     |                                                
  |  |     +-------------------- Uniomerus_carolinianus **     
  |  |     |                                                
  |  |     |                 +-- Quadrula_quadrula **          
  |  |     +-----------------+                              
  |  |     |                 +-- Quadrula_verrucosa **         
  |  |     |                                                
  |  |     |                 +-- Sinohyriopsis_cumingii **     
  |  |     +-----------------+                              
  |  |     |                 +-- Sinohyriopsis_schlegelii **   
  |  |     |                                                
  |  |     |                 +-- Potomida_littoralis **        
  |  |     |           +-----+                              
  |  |     |           |     +-- Pronodularia_japanensis **    
  |  |     +-----------+                                    
  |  |     |           |  +----- Lamprotula_leaii **           
  |  |     |           +--+                                 
  |  |     |              |  +-- Lamprotula_caveata **         
  |  |     |              +--+                              
  |  |     |                 +-- Lamprotula_cornuumlunae **    
  |  |     |                                                
  |  |     |           +-------- Sinosolenaia_carinata **      
  +--+     |           |                                    
     |     |           |  +----- Gonidea_angulata **           
     |     |           +--+                                 
     |     |           |  |  +-- Leguminaia_wheatleyi **       
     |     |           |  +--+                              
     |     |           |     +-- Microcondylaea_bonellii **    
     |     +-----------+                                    
     |     |           |  +----- Ptychorhynchus_pfisteri **    
     |     |           +--+                                 
     |     |           |  |  +-- Parvasolenaia_rivularis **    
     |     |           |  +--+                              
     |     |           |     +-- Koreosolenaia_sitgyensis **   
     |     |           |                                    
     |     |           |  +----- Sinosolenaia_recognita **     
     |     |           +--+                                 
     |     |              |  +-- Sinosolenaia_iridinea **      
     |     |              +--+                              
     |     |                 +-- Sinosolenaia_oleivora **      
     |     |                                                
     |     |              +----- Indopseudodon_salwenianus **  
     |     |        +-----+                                 
     |     |        |     |  +-- Indopseudodon_indawgyiensis **
     |     |        |     +--+                              
     |     |        |        +-- Indopseudodon_bogani **       
     |     |        |                                       
     |     |        |  +-------- Nyeinchanconcha_nyeinchani ** 
     |     |        |  |                                    
     |     |        |  +-------- Thaiconcha_callifera **       
     |     +--------+  |                                    
     |     |        |  +-------- Sundadontina_cumingii **      
     |     |        |  |                                    
     |     |        |  +-------- Sundadontina_tumida[1] **     
     |     |        |  |                                    
     |     |        |  +-------- Sundadontina_tumida[2] **     
     |     |        |  |                                    
     |     |        |  |  +----- Pilsbryoconcha_exilis[2] **   
     |     |        +--+--+                                 
     |     |           |  |  +-- Pilsbryoconcha_carinifera **  
     |     |           |  +--+                              
     |     |           |     +-- Pilsbryoconcha_exilis[1] **   
     |     |           |                                    
     |     |           |  +----- Pseudodon_vagulus **          
     |     |           +--+                                 
     |     |           |  |  +-- Pseudodon_mekongi **          
     |     |           |  +--+                              
     |     |           |     +-- Pseudodon_vondembuschianus ** 
     |     |           |                                    
     |     |           |  +----- Bineurus_loeiensis **         
     |     |           +--+                                 
     |     |              |  +-- Bineurus_mouhotii **          
     |     |              +--+                              
     |     |                 +-- Bineurus_exilis **            
     |     |                                                
     |     |     +-------------- Amblema_plicata **            
     |     |     |                                          
     |     |     +-------------- Toxolasma_parvum **           
     |     |     |                                          
     |     |     |           +-- Obliquaria_reflexa **         
     |     |     +-----------+                              
     |     |     |           +-- Plectomerus_dombeyanus **     
     |     |     |                                          
     |     |     |  +----------- Elliptoideus_sloatianus **    
     |     |     +--+                                       
     |     |     |  |  +-------- Elliptio_complanata **        
     |     |     |  +--+                                    
     |     |     |     |  +----- Eurynia_dilatata **           
     |     +-----+     +--+                                 
     |     |     |        |  +-- Fusconaia_flava **            
     |     |     |        +--+                              
     |     |     |           +-- Pleurobema_sintoxia **        
     |     |     |                                          
     |     |     |     +-------- Ellipsaria_lineolata **       
     |     |     |     |                                    
     |     |     |     +-------- Potamilus_leptodon **         
     |     |     |     |                                    
     |     |     |     +-------- Medionidus_conradicus **      
     |     |     |     |                                    
     +-----+     |     +-------- Potamilus_alatus **           
           |     |     |                                    
           |     |     +-------- Ptychobranchus_subtentus **   
           |     +-----+                                    
           |           +-------- Truncilla_truncata **         
           |           |                                    
           |           +-------- Venustaconcha_ellipsiformis **
           |           |                                    
           |           +-------- Cambarunio_iris **            
           |           |                                    
           |           +-------- Leaunio_lienosus **           
           |           |                                    
           |           |  +----- Ortmanniana_ligamentina **    
           |           +--+                                 
           |              |  +-- Lampsilis_cardium **          
           |              +--+                              
           |                 +-- Lampsilis_ornata **           
           |                                                
           |     +-------------- Hyriopsis_desowitzi **        
           |     |                                          
           |     +-------------- Hyriopsis_myersiana **        
           |     |                                          
           |     +-------------- Hyriopsis_panhai **           
           |     |                                          
           |  +--+           +-- Hyriopsis_bialata **          
           |  |  +-----------+                              
           |  |  |           +-- Hyriopsis_khoratensis **      
           |  |  |                                          
           |  |  |  +----------- Rectidens_sumatrensis **      
           |  |  |  |                                       
           |  |  +--+        +-- Ensidens_telus **             
           |  |     |  +-----+                              
           |  |     |  |     +-- Ensidens_ingallsianus **      
           |  |     +--+                                    
           |  |        |  +----- Ensidens_sagittarius **       
           |  |        +--+                                 
           +--+           |  +-- Ensidens_spiculus **          
           |  |           +--+                              
           |  |              +-- Ensidens_jaculus **           
           |  |                                             
           |  |     +----------- Physunio_modelli **           
           |  |     |                                       
           |  |     +----------- Lens **                       
           |  |     |                                       
           |  |     +----------- Physunio_superbus **          
           |  |     |                                       
           |  |     |  +-------- Yaukthwa_inlenensis **        
           |  |     +--+                                    
           |  +-----+  |  +----- Yaukthwa_panhai **            
           |        |  +--+                                 
           |        |     |  +-- Yaukthwa_nesemanni **         
           |        |     +--+                              
           |        |        +-- Yaukthwa_rectangularis **     
           |        |                                       
           |        |        +-- Lens_comptus **               
           |        |  +-----+                              
           |        |  |     +-- Lens_novoselovi **            
           |        +--+                                    
           |           |  +----- Lens_eximius **               
           |           +--+                                 
           |              |  +-- Lens_contradens **            
           |              +--+                              
           |                 +-- Lens_micropterus **           
           |                                                
           |  +----------------- Acuticosta_chinensis **       
           |  |                                             
           |  +----------------- Alasmidonta_marginata **      
           |  |                                             
           |  +----------------- Anodonta_anatina **           
           |  |                                             
           |  +----------------- Anemina_arcaeformis **        
           |  |                                             
           |  +----------------- Anodonta_cygnea **            
           |  |                                             
           |  +----------------- Cristaria_plicata **          
           |  |                                             
           |  +----------------- Anodonta_nuttalliana **       
           |  |                                             
           |  +----------------- Lasmigona_costata **          
           |  |                                             
           |  +----------------- Lepidodesma_languilati **     
           |  |                                             
           |  +----------------- Pseudanodonta_complanata **   
           |  |                                             
           |  +----------------- Pyganodon_grandis **          
           |  |                                             
           |  +----------------- Simpsonaias_ambigua **        
           |  |                                             
           |  +----------------- Strophitus_undulatus **       
           |  |                                             
           |  |              +-- Schistodesmus_lampreyanus **  
           |  +--------------+                              
           |  |              +-- Schistodesmus_spinosus **     
           |  |                                             
           |  |              +-- Utterbackia_imbecillis **     
           |  +--------------+                              
           |  |              +-- Utterbackia_peninsularis **   
           |  |                                             
           |  |        +-------- Lanceolaria_gladiola **       
           |  +--------+                                    
           |  |        |  +----- Lanceolaria_grayii **         
           |  |        +--+                                 
           +--+           |  +-- Lanceolaria_lanceolata **     
              |           +--+                              
              |              +-- Lanceolaria_triformis **      
              |                                             
              |        +-------- Unio_tumidus **               
              +--------+                                    
              |        |  +----- Unio_crassus **               
              |        +--+                                 
              |           |  +-- Unio_delphinus **             
              |           +--+                              
              |              +-- Unio_pictorum **              
              |                                             
              |        +-------- Aculamprotula_tortuosa **     
              |        |                                    
              +--------+     +-- Aculamprotula_coreana **      
              |        |  +--+                              
              |        |  |  +-- Aculamprotula_scripta **      
              |        +--+                                 
              |           |  +-- Aculamprotula_fibrosa **      
              |           +--+                              
              |              +-- Aculamprotula_tientsinensis **
              |                                             
              |        +-------- Cuneopsis_heudei **           
              |        |                                    
              |        +-------- Cuneopsis_celtiformis **      
              |        |                                    
              +--------+-------- Tchangsinaia_piscicula **     
              |        |                                    
              |        +-------- Cuneopsis_rufescens **        
              |        |                                    
              |        |  +----- Nodularia_douglasiae[2] **    
              |        +--+                                 
              |           |  +-- Nodularia_douglasiae[1] **    
              |           +--+                              
              |              +-- Nodularia_nuxpersicae **      
              |                                             
              |  +-------------- Pletholophus_tenuis **        
              |  |                                          
              +--+           +-- Sinanodonta_angula **         
                 |  +--------+                              
                 |  |        +-- Sinanodonta_lucida **         
                 +--+                                       
                    |  +-------- Sinanodonta_woodiana[1] **    
                    +--+                                    
                       |  +----- Sinanodonta_jourdyi **        
                       +--+                                 
                          |  +-- Sinanodonta_elliptica **      
                          +--+                              
                             +-- Sinanodonta_woodiana[2] **    

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
162 162 155 4 39 13 1 173.0
                                         +-------------------- Gibbosula_rochechouartii     
                                         |                                                  
                                         |  +----------------- Gibbosula_laosensis          
  +--------------------------------------+  |                                               
  |                                      |  |        +-------- Cumberlandia_monodonta       
  |                                      |  |  +-----+                                      
  |                                      +--+  |     |  +----- Pseudunio_marocanus          
  |                                         |  |     +--+                                   
  |                                         |  |        |  +-- Pseudunio_auricularius       
  |                                         |  |        +--+                                
  |                                         +--+           +-- Pseudunio_homsensis          
  |                                            |                                            
  |                                            |           +-- Margaritifera_dahurica       
  |                                            |  +--------+                                
  |                                            |  |        +-- Margaritifera_margaritifera  
  |                                            +--+                                         
  |                                               |  +-------- Margaritifera_falcata        
  |                                               |  |                                      
  |                                               +--+     +-- Margaritifera_hembeli        
  |                                                  |  +--+                                
  |                                                  |  |  +-- Margaritifera_marrianae      
  |                                                  +--+                                   
  |                                                     |  +-- Margaritifera_laevis         
  |                                                     +--+                                
  |                                                        +-- Margaritifera_middendorffi   
  |                                                                                         
  |                                                     +----- Trapezidens_exolescens **       
  |                                               +-----+                                   
  |                                               |     |  +-- Trapezidens_angustior **        
  |                                               |     +--+                                
  |                                               |        +-- Trapezidens_dolichorhynchus **  
  |                                               |                                         
  |                                +--------------+        +-- Lamellidens_corrianus **        
  |                                |              |  +-----+                                
  |                                |              |  |     +-- Lamellidens_savadiensis **      
  |                                |              |  |                                      
  |                                |              |  |  +----- Lamellidens_lamellatus **       
  |                                |              +--+  |                                   
  |                                |                 |  +----- Lamellidens_jenkinsianus **     
  |                                |                 |  |                                   
  |                                |                 |  |  +-- Lamellidens_marginalis **       
  |                                |                 +--+--+                                
  |                                |                    |  +-- Lamellidens_ferrugineus **      
  |                                |                    |                                   
  |  +-----------------------------+                    |  +-- Lamellidens_mainwaringi[1] **   
  |  |                             |                    |  |                                
  |  |                             |                    +--+-- Lamellidens_mainwaringi[2] **   
  |  |                             |                       |                                
  |  |                             |                       +-- Lamellidens_generosus **        
--+  |                             |                                                        
  |  |                             |                    +----- Parreysia_keralaensis **        
  |  |                             |  +-----------------+                                   
  |  |                             |  |                 |  +-- Parreysia_nagpoorensis **       
  |  |                             |  |                 +--+                                
  |  |                             |  |                    +-- Parreysia_plagiosoma **         
  |  |                             |  |                                                     
  |  |                             |  |                 +----- Leoparreysia_canefrii **        
  |  |                             +--+  +--------------+                                   
  |  |                                |  |              |  +-- Leoparreysia_tavoyensis **      
  |  |                                |  |              +--+                                
  |  |                                |  |                 +-- Leoparreysia_olivacea **        
  |  |                                |  |                                                  
  |  |                                |  |        +----------- Coelatura_aegyptiaca[2] **      
  |  |                                |  |        |                                         
  |  |                                +--+  +-----+        +-- Coelatura_aegyptiaca[1] **      
  |  |                                   |  |     |  +-----+                                
  |  |                                   |  |     |  |     +-- Nitia_teretiuscula **           
  |  |                                   |  |     +--+                                      
  |  |                                   |  |        |  +----- Coelatura_kunenensis **         
  |  |                                   |  |        +--+                                   
  |  |                                   |  |           |  +-- Coelatura_gabonensis **         
  |  |                                   |  |           +--+                                
  |  |                                   +--+              +-- Prisodontopsis_mweruensis **    
  |  |                                      |                                               
  |  |                                      |        +-------- Indonaia_shurtleffiana **       
  |  |                                      |        |                                      
  |  |                                      |  +-----+  +----- Indonaia_bonneaudii **          
  |  |                                      |  |     |  |                                   
  |  |                                      |  |     +--+----- Indonaia_rugosa **              
  |  |                                      |  |        |                                   
  |  |                                      |  |        |  +-- Indonaia_andersoniana **        
  |  |                                      +--+        +--+                                
  |  |                                         |           +-- Indonaia_caerulea **            
  |  |                                         |                                            
  |  |                                         |  +----------- Indochinella_pugio **           
  |  |                                         |  |                                         
  |  |                                         |  |     +----- Scabies_phaselus **             
  |  |                                         +--+  +--+                                   
  |  |                                            |  |  |  +-- Scabies_humilis **              
  |  |                                            |  |  +--+                                
  |  |                                            +--+     +-- Scabies_mandarinus **           
  |  |                                               |                                      
  |  |                                               |  +----- Radiatula_mouhoti **            
  |  |                                               +--+                                   
  |  |                                                  |  +-- Radiatula_myitkyinae **         
  |  |                                                  +--+                                
  +--+                                                     +-- Radiatula_chaudhurii **         
     |                                                                                      
     |                                                  +----- Indopseudodon_indawgyiensis **  
     |                                   +--------------+                                   
     |                                   |              |  +-- Indopseudodon_bogani **         
     |                                   |              +--+                                
     |                                   |                 +-- Indopseudodon_salwenianus **    
     |                                   |                                                  
     |                                   |           +-------- Pilsbryoconcha_exilis[3] **     
     |  +--------------------------------+  +--------+                                      
     |  |                                |  |        |  +----- Pilsbryoconcha_exilis[2] **     
     |  |                                |  |        +--+                                   
     |  |                                |  |           |  +-- Pilsbryoconcha_carinifera **    
     |  |                                |  |           +--+                                
     |  |                                |  |              +-- Pilsbryoconcha_exilis[1] **     
     |  |                                |  |                                               
     |  |                                +--+     +----------- Sundadontina_tumida **          
     |  |                                   |  +--+                                         
     |  |                                   |  |  |  +-------- Pseudodon_vagulus **            
     |  |                                   |  |  +--+                                      
     |  |                                   |  |     |  +----- Pseudodon_vondembuschianus[1] **
     |  |                                   |  |     +--+                                   
     |  |                                   |  |        |  +-- Pseudodon_mekongi **            
     |  |                                   +--+        +--+                                
     |  |                                      |           +-- Pseudodon_vondembuschianus[2] **
     |  |                                      |                                            
     |  |                                      |           +-- Nyeinchanconcha_nyeinchani **   
     |  |                                      |  +--------+                                
     |  |                                      |  |        +-- Sundadontina_cumingii **        
     |  |                                      +--+                                         
     |  |                                         |  +-------- Thaiconcha_callifera **         
     |  |                                         +--+                                      
     |  |                                            |  +----- Bineurus_loeiensis **           
     |  |                                            +--+                                   
     |  |                                               |  +-- Bineurus_mouhotii **            
     |  |                                               +--+                                
     |  |                                                  +-- Bineurus_exilis **              
     |  |                                                                                   
     |  |                                                  +-- Ensidens_telus **               
     |  |                                            +-----+                                
     |  |                                            |     +-- Ensidens_ingallsianus **        
     |  |                                      +-----+                                      
     |  |                                      |     |  +----- Ensidens_sagittarius **         
     |  |                                      |     +--+                                   
     |  |                                      |        |  +-- Ensidens_spiculus **            
     |  |                                   +--+        +--+                                
     |  |                                   |  |           +-- Ensidens_jaculus **             
     |  |                                   |  |                                            
     |  |                                   |  |  +----------- Hyriopsis_desowitzi **          
     |  |                                   |  |  |                                         
     |  |                                   |  +--+  +-------- Rectidens_sumatrensis **        
     |  |                                   |     |  |                                      
     |  |                                   |     +--+     +-- Hyriopsis_panhai **             
     +--+                                   |        |  +--+                                
        |                                   |        |  |  +-- Hyriopsis_khoratensis **        
        |  +--------------------------------+        +--+                                   
        |  |                                |           |  +-- Hyriopsis_bialata **            
        |  |                                |           +--+                                
        |  |                                |              +-- Hyriopsis_myersiana **          
        |  |                                |                                               
        |  |                                |           +----- Physunio **                     
        |  |                                |  +--------+                                   
        |  |                                |  |        |  +-- Physunio_superbus **            
        |  |                                |  |        +--+                                
        |  |                                |  |           +-- Physunio_modelli **             
        |  |                                |  |                                            
        |  |                                +--+     +-------- Yaukthwa_inlenensis **          
        |  |                                   |  +--+                                      
        |  |                                   |  |  |  +----- Yaukthwa_panhai **              
        |  |                                   |  |  +--+                                   
        |  |                                   |  |     |  +-- Yaukthwa_nesemanni **           
        |  |                                   |  |     +--+                                
        |  |                                   +--+        +-- Yaukthwa_rectangularis **       
        |  |                                      |                                         
        |  |                                      |        +-- Lens_contradens **              
        |  |                                      |  +-----+                                
        |  |                                      |  |     +-- Lens_micropterus **             
        |  |                                      +--+                                      
        |  |                                         |  +----- Lens_eximius **                 
        |  |                                         +--+                                   
        |  |                                            |  +-- Lens_novoselovi **              
        |  |                                            +--+                                
        |  |                                               +-- Lens_comptus **                 
        |  |                                                                                
        |  |                                            +----- Chamberlainia_somsakpanhai **   
        |  |     +--------------------------------------+                                   
        |  |     |                                      |  +-- Sinohyriopsis_cumingii **       
        |  |     |                                      +--+                                
        |  |     |                                         +-- Sinohyriopsis_schlegelii **     
        |  |     |                                                                          
        |  |     |                                         +-- Potomida_littoralis **          
        |  |     |                                   +-----+                                
        |  |     |                                   |     +-- Pronodularia_japanensis **      
        |  |     |                             +-----+                                      
        |  |     |                             |     |  +----- Lamprotula_leaii **             
        |  |     |                             |     +--+                                   
        |  |     |                             |        |  +-- Lamprotula_caveata **           
        +--+     |                             |        +--+                                
           |     |                             |           +-- Lamprotula_cornuumlunae **      
           |  +--+  +--------------------------+                                            
           |  |  |  |                          |        +----- Sinosolenaia_recognita **       
           |  |  |  |                          |  +-----+                                   
           |  |  |  |                          |  |     |  +-- Sinosolenaia_oleivora **        
           |  |  |  |                          |  |     +--+                                
           |  |  |  |                          |  |        +-- Sinosolenaia_iridinea **        
           |  |  |  |                          +--+                                         
           |  |  |  |                             |     +----- Gonidea_angulata **             
           |  |  |  |                             |  +--+                                   
           |  |  |  |                             |  |  |  +-- Leguminaia_wheatleyi **         
           |  |  |  |                             |  |  +--+                                
           |  |  |  |                             +--+     +-- Microcondylaea_bonellii **      
           |  |  |  |                                |                                      
           |  |  |  |                                |  +----- Ptychorhynchus_pfisteri **      
           |  |  +--+                                +--+                                   
           |  |     |                                   |  +-- Parvasolenaia_rivularis **      
           |  |     |                                   +--+                                
           |  |     |                                      +-- Koreosolenaia_sitgyensis **     
           |  |     |                                                                       
           |  |     |                                      +-- Megalonaias_nervosa **          
           |  |     |                                   +--+                                
           |  |     |                                   |  +-- Uniomerus_carolinianus **       
           |  |     |  +--------------------------------+                                   
           |  |     |  |                                |  +-- Quadrula_quadrula **            
           |  |     |  |                                +--+                                
           |  |     |  |                                   +-- Quadrula_verrucosa **           
           |  |     |  |                                                                    
           |  |     |  |                             +-------- Elliptoideus_sloatianus **      
           |  |     +--+                             |                                      
           |  |        |  +--------------------------+     +-- Elliptio_complanata **          
           |  |        |  |                          |  +--+                                
           |  |        |  |                          |  |  +-- Eurynia_dilatata **             
           |  |        |  |                          +--+                                   
           |  |        |  |                             |  +-- Fusconaia_flava **              
           |  |        +--+                             +--+                                
           |  |           |                                +-- Pleurobema_sintoxia **          
           |  |           |                                                                 
           |  |           |  +-------------------------------- Amblema_plicata **              
           |  |           |  |                                                              
           |  |           +--+  +----------------------------- Toxolasma_parvum **             
           |  |              |  |                                                           
           +--+              +--+                          +-- Obliquaria_reflexa **           
              |                 |  +-----------------------+                                
              |                 |  |                       +-- Plectomerus_dombeyanus **       
              |                 +--+                                                        
              |                    |  +----------------------- Potamilus_leptodon **           
              |                    |  |                                                     
              |                    +--+  +-------------------- Potamilus_alatus **             
              |                       |  |                                                  
              |                       |  |                 +-- Medionidus_conradicus **        
              |                       +--+  +--------------+                                
              |                          |  |              +-- Ptychobranchus_subtentus **     
              |                          |  |                                               
              |                          +--+              +-- Ellipsaria_lineolata **         
              |                             |  +-----------+                                
              |                             |  |           +-- Truncilla_truncata **           
              |                             +--+                                            
              |                                |  +----------- Leaunio_lienosus **             
              |                                |  |                                         
              |                                +--+        +-- Venustaconcha_ellipsiformis **  
              |                                   |  +-----+                                
              |                                   |  |     +-- Cambarunio_iris **              
              |                                   +--+                                      
              |                                      |  +----- Ortmanniana_ligamentina **      
              |                                      +--+                                   
              |                                         |  +-- Lampsilis_cardium **            
              |                                         +--+                                
              |                                            +-- Lampsilis_ornata **             
              |                                                                             
              |           +----------------------------------- Acuticosta_chinensis **         
              |           |                                                                 
              |           |                          +-------- Aculamprotula_tortuosa **       
              |           |                          |                                      
              |           |              +-----------+     +-- Aculamprotula_coreana **        
              |           |              |           |  +--+                                
              |           |              |           |  |  +-- Aculamprotula_scripta **        
              |           |              |           +--+                                   
              |           |              |              |  +-- Aculamprotula_fibrosa **        
              |           |              |              +--+                                
              |           |              |                 +-- Aculamprotula_tientsinensis **  
              |           |  +-----------+                                                  
              |           |  |           |           +-------- Unio_tumidus **                 
              +-----------+  |           |  +--------+                                      
                          |  |           |  |        |  +----- Unio_crassus **                 
                          |  |           |  |        +--+                                   
                          |  |           |  |           |  +-- Unio_delphinus **               
                          |  |           |  |           +--+                                
                          |  |           +--+              +-- Unio_pictorum **                
                          |  |              |                                               
                          |  |              |           +----- Nodularia_douglasiae[1] **      
                          |  |              |  +--------+                                   
                          |  |              |  |        |  +-- Nodularia_douglasiae[2] **      
                          |  |              |  |        +--+                                
                          |  |              +--+           +-- Nodularia_nuxpersicae **        
                          |  |                 |                                            
                          +--+                 |  +----------- Schistodesmus_lampreyanus **    
                             |                 +--+                                         
                             |                    |  +-------- Tchangsinaia_piscicula **       
                             |                    +--+                                      
                             |                       |  +----- Cuneopsis_rufescens **          
                             |                       +--+                                   
                             |                          |  +-- Cuneopsis_heudei **             
                             |                          +--+                                
                             |                             +-- Cuneopsis_celtiformis **        
                             |                                                              
                             |  +----------------------------- Lepidodesma_languilati **       
                             |  |                                                           
                             |  |                    +-------- Lanceolaria_gladiola **         
                             |  |  +-----------------+                                      
                             |  |  |                 |  +----- Lanceolaria_grayii **           
                             |  |  |                 +--+                                   
                             +--+  |                    |  +-- Lanceolaria_lanceolata **       
                                |  |                    +--+                                
                                |  |                       +-- Lanceolaria_triformis **        
                                |  |                                                        
                                |  |     +-------------------- Anemina_arcaeformis **          
                                |  |     |                                                  
                                |  |  +--+  +----------------- Cristaria_plicata **            
                                +--+  |  |  |                                               
                                   |  |  +--+  +-------------- Pletholophus_tenuis **          
                                   |  |     |  |                                            
                                   |  |     +--+           +-- Sinanodonta_angula **           
                                   |  |        |  +--------+                                
                                   |  |        |  |        +-- Sinanodonta_lucida **           
                                   |  |        +--+                                         
                                   |  |           |  +-------- Sinanodonta_woodiana[2] **      
                                   |  |           +--+                                      
                                   |  |              |  +----- Sinanodonta_jourdyi **          
                                   +--+              +--+                                   
                                      |                 |  +-- Sinanodonta_elliptica **        
                                      |                 +--+                                
                                      |                    +-- Sinanodonta_woodiana[1] **      
                                      |                                                     
                                      |                    +-- Alasmidonta_marginata **        
                                      |           +--------+                                
                                      |           |        +-- Lasmigona_costata **            
                                      |           |                                         
                                      |           |     +----- Anodonta_anatina **             
                                      |           +-----+                                   
                                      |           |     |  +-- Anodonta_cygnea **              
                                      +-----------+     +--+                                
                                                  |        +-- Anodonta_nuttalliana **         
                                                  |                                         
                                                  |     +----- Pseudanodonta_complanata **     
                                                  |  +--+                                   
                                                  |  |  |  +-- Simpsonaias_ambigua **          
                                                  |  |  +--+                                
                                                  +--+     +-- Strophitus_undulatus **         
                                                     |                                      
                                                     |  +----- Pyganodon_grandis **            
                                                     +--+                                   
                                                        |  +-- Utterbackia_imbecillis **       
                                                        +--+                                
                                                           +-- Utterbackia_peninsularis **     

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
131 131 72 4 0 0 0 148.0
                          +----- Trapezidens_exolescens **       
                 +--------+                                   
                 |        |  +-- Trapezidens_dolichorhynchus **  
                 |        +--+                                
     +-----------+           +-- Trapezidens_angustior **        
     |           |                                            
     |           |  +----------- Lamellidens_lamellatus **       
     |           |  |                                         
     |           +--+  +-------- Lamellidens_jenkinsianus **     
     |              |  |                                      
     |              |  |  +----- Lamellidens_generosus **        
     |              +--+  |                                   
     |                 |  +----- Lamellidens_mainwaringi **      
     |                 |  |                                   
     |                 +--+  +-- Lamellidens_ferrugineus **      
     |                    +--+                                
     |                    |  +-- Lamellidens_marginalis **       
     |                    |                                   
  +--+                    |  +-- Lamellidens_savadiensis **      
  |  |                    +--+                                
  |  |                       +-- Lamellidens_corrianus[1] **     
  |  |                                                        
  |  |                 +-------- Parreysia_keralaensis **        
  |  |  +--------------+                                      
  |  |  |              |  +----- Parreysia_plagiosoma **         
  |  |  |              +--+                                   
  |  |  |                 |  +-- Parreysia_corrugata **          
  |  |  |                 +--+                                
  |  |  |                    +-- Parreysia_nagpoorensis **       
  |  |  |                                                     
  |  |  |              +-------- Leoparreysia_olivacea **        
  |  |  |  +-----------+                                      
  |  |  |  |           |  +----- Leoparreysia_tavoyensis[2] **   
  |  +--+  |           +--+                                   
  |     |  |              |  +-- Leoparreysia_canefrii **        
  |     |  |              +--+                                
  |     |  |                 +-- Leoparreysia_tavoyensis[1] **   
  |     |  |                                                  
  |     |  |                 +-- Nitia_teretiuscula **           
  |     |  |                 |                                
  |     |  |                 +-- Coelatura_gabonensis **         
  |     |  |                 |                                
  |     |  |                 +-- Prisodontopsis_mweruensis **    
  |     +--+                 |                                
  |        |                 +-- Coelatura_kunenensis **         
  |        |                 |                                
  |        |  +--------------+-- Coelatura_aegyptiaca[1] **      
  |        |  |              |                                
  |        |  |              +-- Coelatura_aegyptiaca[2] **      
  |        |  |              |                                
  |        |  |              +-- Prisodontopsis_aviculaeformis **
  |        |  |              |                                
  |        |  |              +-- Coelatura_luapulaensis **       
  |        |  |              |                                
  |        |  |              +-- Coelatura_choziensis **         
  |        +--+                                               
  |           |        +-------- Indonaia_shurtleffiana **       
  |           |        |                                      
  |           |  +-----+  +----- Indonaia_rugosa **              
  |           |  |     |  |                                   
  |           |  |     |  |  +-- Indonaia_andersoniana **        
  |           |  |     +--+--+                                
  |           |  |        |  +-- Indonaia_caerulea **            
  |           |  |        |                                   
  |           +--+        |  +-- Indonaia_bonneaudii **          
  |              |        +--+                                
  |              |           +-- Lamellidens_corrianus[2] **     
  |              |                                            
  |              |  +----------- Indochinella_pugio **           
  |              |  |                                         
  |              |  |        +-- Scabies_humilis **              
  |              +--+        |                                
  |                 |  +-----+-- Scabies_phaselus **             
--+                 |  |     |                                
  |                 +--+     +-- Scabies_mandarinus **           
  |                    |                                      
  |                    |  +----- Radiatula_mouhoti **            
  |                    +--+                                   
  |                       |  +-- Radiatula_myitkyinae **         
  |                       +--+                                
  |                          +-- Radiatula_chaudhurii **         
  |                                                           
  |                       +----- Indopseudodon_indawgyiensis **  
  |           +-----------+                                   
  |           |           |  +-- Indopseudodon_salwenianus **    
  |           |           +--+                                
  |           |              +-- Indopseudodon_bogani **         
  |           |                                               
  |           |              +-- Nyeinchanconcha_nyeinchani **   
  |           |  +-----------+                                
  |     +-----+  |           +-- Sundadontina_cumingii **        
  |     |     |  |                                            
  |     |     |  |        +----- Bineurus_exilis **              
  |     |     |  +--------+                                   
  |     |     |  |        |  +-- Bineurus_mouhotii **            
  |     |     |  |        +--+                                
  |     |     |  |           +-- Bineurus_loeiensis **           
  |     |     |  |                                            
  |     |     +--+     +-------- Sundadontina_tumida **          
  |     |        +-----+                                      
  |     |        |     |  +----- Pseudodon_vagulus **            
  |     |        |     +--+                                   
  |     |        |        |  +-- Pseudodon_mekongi **            
  |     |        |        +--+                                
  |     |        |           +-- Pseudodon_vondembuschianus **   
  |     |        |                                            
  |     |        |  +----------- Thaiconcha_callifera **         
  |     |        +--+                                         
  |     |           |  +-------- Pilsbryoconcha_exilis[3] **     
  |     |           +--+                                      
  |     |              |  +----- Pilsbryoconcha_exilis[2] **     
  |     |              +--+                                   
  |     |                 |  +-- Pilsbryoconcha_exilis[1] **     
  |     |                 +--+                                
  |     |                    +-- Pilsbryoconcha_carinifera **    
  |     |                                                     
  |     |                    +-- Ensidens_telus **               
  |     |              +-----+                                
  |     |              |     +-- Ensidens_ingallsianus **        
  |     |           +--+                                      
  |     |           |  |  +----- Ensidens_jaculus **             
  |     |           |  +--+                                   
  |     |           |     |  +-- Ensidens_spiculus **            
  |     |           |     +--+                                
  |     |     +-----+        +-- Ensidens_sagittarius **         
  |     |     |     |                                         
  |     |     |     |  +-------- Hyriopsis_desowitzi **          
  |     |     |     |  |                                      
  |     |     |     |  +-------- Rectidens_sumatrensis **        
  +-----+     |     +--+                                      
        |     |        |  +----- Hyriopsis_panhai **             
        |     |        |  |                                   
        |     |        +--+  +-- Hyriopsis_myersiana **          
        |     |           |  |                                
        |  +--+           +--+-- Hyriopsis_bialata **            
        |  |  |              |                                
        |  |  |              +-- Hyriopsis_khoratensis **        
        |  |  |                                               
        |  |  |           +----- Physunio **                     
        |  |  |  +--------+                                   
        |  |  |  |        |  +-- Physunio_superbus **            
        |  |  |  |        +--+                                
        |  |  |  |           +-- Physunio_modelli **             
        |  |  |  |                                            
        |  |  +--+     +-------- Yaukthwa_inlenensis **          
        |  |     |  +--+                                      
        |  |     |  |  |  +----- Yaukthwa_panhai **              
        |  |     |  |  +--+                                   
        |  |     |  |     |  +-- Yaukthwa_nesemanni **           
        |  |     |  |     +--+                                
        |  |     +--+        +-- Yaukthwa_rectangularis **       
        |  |        |                                         
        |  |        |        +-- Lens_contradens **              
        |  |        |  +-----+                                
        |  |        |  |     +-- Lens_micropterus **             
        |  |        +--+                                      
        |  |           |  +----- Lens_eximius **                 
        |  |           +--+                                   
        |  |              |  +-- Lens_comptus **                 
        |  |              +--+                                
        |  |                 +-- Lens_novoselovi **              
        |  |                                                  
        |  |                 +-- Amblema_plicata **              
        |  |                 |                                
        |  |                 +-- Ortmanniana_ligamentina **      
        |  |                 |                                
        |  |                 +-- Lampsilis_cardium **            
        |  |                 |                                
        |  |                 +-- Cambarunio_iris **              
        +--+                 |                                
           |              +--+-- Elliptio_complanata **          
           |              |  |                                
           |              |  +-- Eurynia_dilatata **             
           |              |  |                                
           |              |  +-- Pleurobema_sintoxia **          
           |              |  |                                
           |              |  +-- Quadrula_quadrula **            
           |              |  |                                
           |              |  +-- Quadrula_verrucosa **           
           |              |                                   
           |              |  +-- Chamberlainia_somsakpanhai **   
           |           +--+  |                                
           |           |  |  +-- Sinohyriopsis_cumingii **       
           |           |  |  |                                
           |           |  |  +-- Lamprotula_caveata **           
           |           |  |  |                                
           |           |  |  +-- Lamprotula_leaii **             
           |           |  |  |                                
           |           |  |  +-- Potomida_littoralis **          
           |           |  |  |                                
           |           |  |  +-- Pronodularia_japanensis **      
           |           |  |  |                                
           |           |  +--+-- Gonidea_angulata **             
           |           |     |                                
           |           |     +-- Leguminaia_wheatleyi **         
           |           |     |                                
           |           |     +-- Leguminaia **                   
           |           |     |                                
           |           |     +-- Microcondylaea_bonellii **      
           |           |     |                                
           |           |     +-- Sinosolenaia_carinata **        
           |           |     |                                
           |           |     +-- Sinosolenaia_oleivora **        
           |           |     |                                
           |           |     +-- Sinosolenaia_recognita **       
           +-----------+                                      
                       |     +-- Unio_crassus **                 
                       |     |                                
                       |     +-- Unio_pictorum **                
                       |     |                                
                       |     +-- Unio_tumidus **                 
                       |     |                                
                       |     +-- Aculamprotula_tortuosa **       
                       |     |                                
                       |     +-- Cuneopsis_heudei **             
                       |  +--+                                
                       |  |  +-- Tchangsinaia_piscicula **       
                       |  |  |                                
                       |  |  +-- Cuneopsis_rufescens **          
                       |  |  |                                
                       |  |  +-- Nodularia_douglasiae **         
                       |  |  |                                
                       |  |  +-- Nodularia_nuxpersicae **        
                       |  |  |                                
                       |  |  +-- Schistodesmus_lampreyanus **    
                       |  |                                   
                       |  |  +-- Alasmidonta_marginata **        
                       |  |  |                                
                       |  |  +-- Anodonta_anatina **             
                       +--+  |                                
                          |  +-- Anodonta_cygnea **              
                          |  |                                
                          |  +-- Anodonta_nuttalliana **         
                          |  |                                
                          |  +-- Lasmigona_costata **            
                          |  |                                
                          |  +-- Pseudanodonta_complanata **     
                          |  |                                
                          |  +-- Pyganodon_grandis **            
                          |  |                                
                          |  +-- Simpsonaias_ambigua **          
                          |  |                                
                          |  +-- Strophitus_undulatus **         
                          +--+                                
                             +-- Sinanodonta_lauta **            
                             |                                
                             +-- Cristaria_plicata **            
                             |                                
                             +-- Cristaria_radiata **            
                             |                                
                             +-- Pletholophus_tenuis **          
                             |                                
                             +-- Sinanodonta_lucida **           
                             |                                
                             +-- Sinanodonta_jourdyi **          
                             |                                
                             +-- Lanceolaria_gladiola **         
                             |                                
                             +-- Lanceolaria_grayii[1] **        
                             |                                
                             +-- Lanceolaria_grayii[2] **        

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
116 116 114 4 25 10 1 134.0
                                   +-- Trapezidens_dolichorhynchus **
                       +-----------+                              
                       |           +-- Trapezidens_exolescens **     
           +-----------+                                          
           |           |  +----------- Lamellidens_corrianus **      
           |           |  |                                       
           |           +--+  +-------- Lamellidens_savadiensis **    
           |              |  |                                    
           |              +--+     +-- Lamellidens_jenkinsianus **   
           |                 |  +--+                              
           |                 |  |  +-- Lamellidens_mainwaringi **    
           |                 +--+                                 
  +--------+                    |  +-- Lamellidens_ferrugineus **    
  |        |                    +--+                              
  |        |                       +-- Lamellidens_marginalis **     
  |        |                                                      
  |        |                       +-- Parreysia_nagpoorensis **     
  |        |  +--------------------+                              
  |        |  |                    +-- Parreysia_plagiosoma **       
  |        |  |                                                   
  |        |  |                    +-- Leoparreysia_tavoyensis **    
  |        |  |  +-----------------+                              
  |        +--+  |                 +-- Leoparreysia_olivacea **      
  |           |  |                                                
  |           |  |     +-------------- Nitia_teretiuscula **         
  |           |  |  +--+                                          
  |           |  |  |  |  +----------- Coelatura_aegyptiaca[2] **    
  |           +--+  |  +--+                                       
  |              |  |     |  +-------- Coelatura_aegyptiaca[1] **    
  |              |  |     +--+                                    
  |              |  |        |  +----- Prisodontopsis_mweruensis **  
  |              |  |        +--+                                 
  |              |  |           |  +-- Coelatura_gabonensis **       
  |              +--+           +--+                              
  |                 |              +-- Coelatura_kunenensis **       
  |                 |                                             
  |                 |        +-------- Indonaia_shurtleffiana **     
  |                 |  +-----+                                    
  |                 |  |     |  +----- Indonaia_andersoniana **      
  |                 |  |     +--+                                 
  |                 |  |        |  +-- Indonaia_rugosa **            
  |                 |  |        +--+                              
  |                 +--+           +-- Indonaia_bonneaudii **        
  |                    |                                          
  |                    |           +-- Indochinella_pugio **         
  |                    |  +--------+                              
  |                    |  |        +-- Scabies_crispata **           
  |                    |  |                                       
  |                    +--+        +-- Radiatula_myitkyinae **       
  |                       |  +-----+                              
  |                       |  |     +-- Radiatula_chaudhurii **       
  |                       +--+                                    
  |                          |  +----- Scabies_mandarinus **         
  |                          +--+                                 
  |                             |  +-- Scabies_humilis **            
  |                             +--+                              
  |                                +-- Scabies_phaselus **           
  |                                                               
  |                                +-- Ensidens_telus **             
  |                          +-----+                              
--+                          |     +-- Ensidens_ingallsianus **      
  |                    +-----+                                    
  |                    |     |  +----- Ensidens_sagittarius **       
  |                    |     +--+                                 
  |                    |        |  +-- Ensidens_spiculus **          
  |                    |        +--+                              
  |                 +--+           +-- Ensidens_jaculus **           
  |                 |  |                                          
  |                 |  |           +-- Hyriopsis_bialata **          
  |                 |  |  +--------+                              
  |                 |  |  |        +-- Hyriopsis_khoratensis **      
  |                 |  +--+                                       
  |                 |     |  +-------- Hyriopsis_panhai **           
  |                 |     +--+                                    
  |                 |        |  +----- Hyriopsis_myersiana **        
  |                 |        +--+                                 
  |  +--------------+           |  +-- Rectidens_sumatrensis **      
  |  |              |           +--+                              
  |  |              |              +-- Hyriopsis_desowitzi **        
  |  |              |                                             
  |  |              |           +----- Physunio **                   
  |  |              |  +--------+                                 
  |  |              |  |        |  +-- Physunio_modelli **           
  |  |              |  |        +--+                              
  |  |              |  |           +-- Physunio_superbus **          
  |  |              |  |                                          
  |  |              +--+     +-------- Yaukthwa_inlenensis **        
  |  |                 |  +--+                                    
  |  |                 |  |  |  +----- Yaukthwa_panhai **            
  |  |                 |  |  +--+                                 
  |  |                 |  |     |  +-- Yaukthwa_rectangularis **     
  |  |                 |  |     +--+                              
  |  |                 +--+        +-- Yaukthwa_nesemanni **         
  |  |                    |                                       
  |  |                    |        +-- Lens_contradens **            
  |  |                    |  +-----+                              
  |  |                    |  |     +-- Lens_micropterus **           
  |  |                    +--+                                    
  |  |                       |  +----- Lens_eximius **               
  |  |                       +--+                                 
  |  |                          |  +-- Lens_comptus **               
  |  |                          +--+                              
  |  |                             +-- Lens_novoselovi **            
  |  |                                                            
  |  |           +-------------------- Indopseudodon_indawgyiensis **
  |  |           |                                                
  |  |           |           +-------- Pilsbryoconcha_exilis[1] **   
  |  |           |  +--------+                                    
  +--+  +--------+  |        |  +----- Pilsbryoconcha_exilis[3] **   
     |  |        |  |        +--+                                 
     |  |        |  |           |  +-- Pilsbryoconcha_exilis[2] **   
     |  |        |  |           +--+                              
     |  |        |  |              +-- Pilsbryoconcha_carinifera **  
     |  |        +--+                                             
     |  |           |        +-------- Sundadontina_tumida **        
     |  |           |  +-----+                                    
     |  |           |  |     |  +----- Pseudodon_vagulus **          
     |  |           |  |     +--+                                 
     |  |           |  |        |  +-- Pseudodon_mekongi **          
     |  |           |  |        +--+                              
     |  |           +--+           +-- Pseudodon_vondembuschianus ** 
     |  |              |                                          
     |  |              |           +-- Nyeinchanconcha_nyeinchani ** 
     |  |              |  +--------+                              
     |  |              |  |        +-- Sundadontina_cumingii **      
     |  |              +--+                                       
     |  |                 |  +-------- Thaiconcha_callifera **       
     |  |                 +--+                                    
     |  |                    |  +----- Bineurus_exilis **            
     |  |                    +--+                                 
     |  |                       |  +-- Bineurus_mouhotii **          
     |  |                       +--+                              
     |  |                          +-- Bineurus_loeiensis **         
     |  |                                                         
     |  |                    +-------- Amblema_plicata **            
     |  |                 +--+                                    
     |  |                 |  |  +----- Ortmanniana_ligamentina **    
     |  |                 |  +--+                                 
     |  |                 |     |  +-- Lampsilis_cardium **          
     |  |                 |     +--+                              
     +--+           +-----+        +-- Cambarunio_iris **            
        |           |     |                                       
        |           |     |        +-- Quadrula_quadrula **          
        |           |     |  +-----+                              
        |           |     |  |     +-- Quadrula_verrucosa **         
        |           |     +--+                                    
        |           |        |  +----- Pleurobema_sintoxia **        
        |           |        +--+                                 
        |           |           |  +-- Eurynia_dilatata **           
        |           |           +--+                              
        |           |              +-- Elliptio_complanata **        
        |  +--------+                                             
        |  |        |              +-- Sinohyriopsis_cumingii **     
        |  |        |        +-----+                              
        |  |        |        |     +-- Chamberlainia_somsakpanhai ** 
        |  |        |        |                                    
        |  |        |  +-----+     +-- Potomida_littoralis **        
        |  |        |  |     |  +--+                              
        |  |        |  |     |  |  +-- Pronodularia_japanensis **    
        |  |        |  |     +--+                                 
        |  |        |  |        |  +-- Lamprotula_caveata **         
        |  |        |  |        +--+                              
        |  |        +--+           +-- Lamprotula_leaii **           
        |  |           |                                          
        |  |           |        +----- Sinosolenaia_carinata **      
        |  |           |  +-----+                                 
        |  |           |  |     |  +-- Sinosolenaia_oleivora **      
        |  |           |  |     +--+                              
        |  |           +--+        +-- Sinosolenaia_recognita **     
        |  |              |                                       
        |  |              |  +-------- Gonidea_angulata **           
        |  |              +--+                                    
        +--+                 |  +----- Microcondylaea_bonellii **    
           |                 +--+                                 
           |                    |  +-- Leguminaia_wheatleyi **       
           |                    +--+                              
           |                       +-- Leguminaia **                 
           |                                                      
           |                    +----- Unio_tumidus **               
           |           +--------+                                 
           |           |        |  +-- Unio_pictorum **              
           |           |        +--+                              
           |           |           +-- Unio_crassus **               
           |  +--------+                                          
           |  |        |  +----------- Aculamprotula_tortuosa **     
           |  |        |  |                                       
           |  |        |  |        +-- Nodularia_douglasiae **       
           |  |        +--+  +-----+                              
           |  |           |  |     +-- Nodularia_nuxpersicae **      
           |  |           |  |                                    
           |  |           +--+     +-- Cuneopsis_rufescens **        
           |  |              |  +--+                              
           |  |              |  |  +-- Cuneopsis_heudei **           
           |  |              +--+                                 
           +--+                 |  +-- Tchangsinaia_piscicula **     
              |                 +--+                              
              |                    +-- Schistodesmus_lampreyanus **  
              |                                                   
              |                    +-- Lanceolaria_grayii **         
              |  +-----------------+                              
              |  |                 +-- Lanceolaria_gladiola **       
              |  |                                                
              |  |                 +-- Cristaria_plicata **          
              |  |        +--------+                              
              |  |        |        +-- Cristaria_radiata **          
              +--+  +-----+                                       
                 |  |     |  +-------- Pletholophus_tenuis **        
                 |  |     +--+                                    
                 |  |        |  +----- Sinanodonta_lucida **         
                 |  |        +--+                                 
                 |  |           |  +-- Sinanodonta_jourdyi **        
                 +--+           +--+                              
                    |              +-- Sinanodonta_lauta **          
                    |                                             
                    |  +-------------- Anodonta_anatina **           
                    |  |                                          
                    |  |        +----- Pseudanodonta_complanata **   
                    +--+  +-----+                                 
                       |  |     |  +-- Anodonta_cygnea **            
                       |  |     +--+                              
                       |  |        +-- Anodonta_nuttalliana **       
                       +--+                                       
                          |        +-- Lasmigona_costata **          
                          |  +-----+                              
                          |  |     +-- Alasmidonta_marginata **      
                          +--+                                    
                             |  +----- Simpsonaias_ambigua **        
                             +--+                                 
                                |  +-- Pyganodon_grandis **          
                                +--+                              
                                   +-- Strophitus_undulatus **       

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
116 139 123 4 25 10 1 151.5
              +-------------------- Chamberlainia_somsakpanhai ** 
              |                                                
  +-----------+  +----------------- Cumberlandia_monodonta     
  |           |  |                                             
  |           |  |  +-------------- Gibbosula_laosensis        
  |           +--+  |                                          
  |              |  |           +-- Pseudunio_auricularius     
  |              |  +-----------+                              
  |              +--+           +-- Pseudunio_marocanus        
  |                 |                                          
  |                 |           +-- Margaritifera_dahurica     
  |                 |  +--------+                              
  |                 |  |        +-- Margaritifera_margaritifera
  |                 +--+                                       
  |                    |  +-------- Margaritifera_falcata      
  |                    +--+                                    
  |                       |  +----- Margaritifera_middendorffi 
  |                       +--+                                 
  |                          |  +-- Margaritifera_laevis       
  |                          +--+                              
  |                             +-- Margaritifera_marrianae    
  |                                                            
  |                             +-- Tamsiella_schroeteriana    
  |                          +--+                              
  |                          |  +-- Etheria_elliptica          
  |                 +--------+                                 
  |                 |        |  +-- Aspatharia_pfeifferiana    
  |                 |        +--+                              
  |                 |           +-- Chambardia_wahlbergi       
  |                 |                                          
  +-----------------+        +----- Diplodon_suavidicus        
  |                 |  +-----+                                 
  |                 |  |     |  +-- Castalia_ambigua           
  |                 |  |     +--+                              
  |                 |  |        +-- Triplodon_corrugatus       
  |                 +--+                                       
  |                    |     +----- Velesunio_ambiguus         
  |                    |  +--+                                 
  |                    |  |  |  +-- Alathyria_jacksoni         
  |                    |  |  +--+                              
  |                    +--+     +-- Alathyria_profuga          
  |                       |                                    
  |                       |  +----- Microdontia_anodontaeformis
  |                       +--+                                 
  |                          |  +-- Alathyria_pertexta         
  |                          +--+                              
  |                             +-- Lortiella_froggatti        
  |                                                            
  |                             +-- Parreysia_nagpoorensis **     
  |                 +-----------+                              
  |                 |           +-- Parreysia_plagiosoma **       
  |                 |                                          
  |        +--------+           +-- Trapezidens_dolichorhynchus **
  |        |        |  +--------+                              
  |        |        |  |        +-- Trapezidens_exolescens **     
  |        |        |  |                                       
  |        |        +--+  +-------- Lamellidens_savadiensis **    
--+        |           |  |                                    
  |        |           |  +-------- Lamellidens_corrianus **      
  |        |           +--+                                    
  |        |              |     +-- Lamellidens_marginalis **     
  |        |              |  +--+                              
  |        |              |  |  +-- Lamellidens_ferrugineus **    
  |  +-----+              +--+                                 
  |  |     |                 |  +-- Lamellidens_jenkinsianus **   
  |  |     |                 +--+                              
  |  |     |                    +-- Lamellidens_mainwaringi **    
  |  |     |                                                   
  |  |     |                    +-- Leoparreysia_olivacea **      
  |  |     |  +-----------------+                              
  |  |     |  |                 +-- Leoparreysia_tavoyensis **    
  |  |     |  |                                                
  |  |     |  |     +-------------- Nitia_teretiuscula **         
  |  |     |  |  +--+                                          
  |  |     |  |  |  |  +----------- Coelatura_aegyptiaca[1] **    
  |  |     +--+  |  +--+                                       
  |  |        |  |     |  +-------- Coelatura_aegyptiaca[2] **    
  |  |        |  |     +--+                                    
  |  |        |  |        |  +----- Coelatura_gabonensis **       
  |  |        |  |        +--+                                 
  |  |        |  |           |  +-- Coelatura_kunenensis **       
  |  |        +--+           +--+                              
  |  |           |              +-- Prisodontopsis_mweruensis **  
  |  |           |                                             
  |  |           |        +-------- Indonaia_shurtleffiana **     
  |  |           |  +-----+                                    
  |  |           |  |     |  +----- Indonaia_andersoniana **      
  |  |           |  |     +--+                                 
  |  |           |  |        |  +-- Indonaia_rugosa **            
  |  |           |  |        +--+                              
  |  |           +--+           +-- Indonaia_bonneaudii **        
  |  |              |                                          
  |  |              |           +-- Indochinella_pugio **         
  |  |              |  +--------+                              
  |  |              |  |        +-- Scabies_crispata **           
  |  |              |  |                                       
  |  |              +--+        +-- Radiatula_myitkyinae **       
  |  |                 |  +-----+                              
  |  |                 |  |     +-- Radiatula_chaudhurii **       
  |  |                 +--+                                    
  |  |                    |  +----- Scabies_mandarinus **         
  |  |                    +--+                                 
  |  |                       |  +-- Scabies_humilis **            
  |  |                       +--+                              
  |  |                          +-- Scabies_phaselus **           
  |  |                                                         
  |  |                       +----- Unio_tumidus **               
  |  |                 +-----+                                 
  |  |                 |     |  +-- Unio_crassus **               
  |  |                 |     +--+                              
  |  |                 |        +-- Unio_pictorum **              
  |  |           +-----+                                       
  +--+           |     |  +-------- Aculamprotula_tortuosa **     
     |           |     |  |                                    
     |           |     |  +-------- Schistodesmus_lampreyanus **  
     |           |     |  |                                    
     |           |     +--+     +-- Nodularia_douglasiae **       
     |           |        +-----+                              
     |           |        |     +-- Nodularia_nuxpersicae **      
     |           |        |                                    
     |           |        |  +----- Cuneopsis_heudei **           
     |           |        +--+                                 
     |           |           |  +-- Tchangsinaia_piscicula **     
     |           |           +--+                              
     |  +--------+              +-- Cuneopsis_rufescens **        
     |  |        |                                             
     |  |        |              +-- Lanceolaria_grayii **         
     |  |        |  +-----------+                              
     |  |        |  |           +-- Lanceolaria_gladiola **       
     |  |        |  |                                          
     |  |        |  |           +-- Cristaria_plicata **          
     |  |        |  |  +--------+                              
     |  |        |  |  |        +-- Cristaria_radiata **          
     |  |        |  +--+                                       
     |  |        |  |  |  +-------- Pletholophus_tenuis **        
     |  |        |  |  +--+                                    
     |  |        |  |     |  +----- Sinanodonta_lucida **         
     |  |        +--+     +--+                                 
     |  |           |        |  +-- Sinanodonta_lauta **          
     |  |           |        +--+                              
     |  |           |           +-- Sinanodonta_jourdyi **        
     |  |           |                                          
     |  |           |     +-------- Anodonta_anatina **           
     |  |           |     |                                    
     |  |           |     +-------- Pseudanodonta_complanata **   
     |  |           |     |                                    
     |  |           |     |     +-- Anodonta_cygnea **            
     |  |           +-----+-----+                              
     |  |                 |     +-- Anodonta_nuttalliana **       
     |  |                 |                                    
     |  |                 |  +----- Pyganodon_grandis **          
     |  |                 |  |                                 
     |  |                 |  +----- Simpsonaias_ambigua **        
     |  |                 +--+                                 
     |  |                    +----- Strophitus_undulatus **       
     |  |                    |                                 
     |  |                    |  +-- Alasmidonta_marginata **      
     |  |                    +--+                              
     |  |                       +-- Lasmigona_costata **          
     |  |                                                      
     |  |                 +-------- Lens_contradens **            
     |  |           +-----+                                    
     +--+           |     |  +----- Lens_eximius **               
        |           |     +--+                                 
        |           |        |  +-- Lens_comptus **               
        |           |        +--+                              
        |        +--+           +-- Lens_novoselovi **            
        |        |  |                                          
        |        |  |        +----- Physunio **                   
        |        |  |  +-----+                                 
        |        |  |  |     |  +-- Physunio_modelli **           
        |        |  |  |     +--+                              
        |        |  +--+        +-- Physunio_superbus **          
        |        |     |                                       
        |        |     |  +-------- Yaukthwa_inlenensis **        
        |        |     +--+                                    
        |        |        |  +----- Yaukthwa_panhai **            
        |  +-----+        +--+                                 
        |  |     |           |  +-- Yaukthwa_rectangularis **     
        |  |     |           +--+                              
        |  |     |              +-- Yaukthwa_nesemanni **         
        |  |     |                                             
        |  |     |           +----- Hyriopsis_panhai **           
        |  |     |           |                                 
        |  |     |  +--------+----- Hyriopsis_khoratensis **      
        |  |     |  |        |                                 
        |  |     |  |        |  +-- Hyriopsis_myersiana **        
        |  |     |  |        +--+                              
        |  |     +--+           +-- Hyriopsis_bialata **          
        |  |        |                                          
        |  |        |  +----------- Hyriopsis_desowitzi **        
        |  |        |  |                                       
        |  |        |  +----------- Rectidens_sumatrensis **      
        |  |        +--+                                       
        |  |           |        +-- Ensidens_telus **             
        |  |           |  +-----+                              
        |  |           |  |     +-- Ensidens_ingallsianus **      
        |  |           +--+                                    
        |  |              |  +----- Ensidens_sagittarius **       
        |  |              +--+                                 
        |  |                 |  +-- Ensidens_spiculus **          
        +--+                 +--+                              
           |                    +-- Ensidens_jaculus **           
           |                                                   
           |              +-------- Amblema_plicata **            
           |           +--+                                    
           |           |  |  +----- Cambarunio_iris **            
           |           |  +--+                                 
           |           |     |  +-- Ortmanniana_ligamentina **    
           |           |     +--+                              
           |  +--------+        +-- Lampsilis_cardium **          
           |  |        |                                       
           |  |        |        +-- Quadrula_quadrula **          
           |  |        |  +-----+                              
           |  |        |  |     +-- Quadrula_verrucosa **         
           |  |        +--+                                    
           |  |           |  +----- Pleurobema_sintoxia **        
           |  |           +--+                                 
           |  |              |  +-- Elliptio_complanata **        
           |  |              +--+                              
           |  |                 +-- Eurynia_dilatata **           
           |  |                                                
           |  |           +-------- Sinohyriopsis_cumingii **     
           |  |           |                                    
           |  |     +-----+     +-- Lamprotula_caveata **         
           +--+     |     |  +--+                              
              |     |     |  |  +-- Lamprotula_leaii **           
              |     |     +--+                                 
              |     |        |  +-- Potomida_littoralis **        
              |  +--+        +--+                              
              |  |  |           +-- Pronodularia_japanensis **    
              |  |  |                                          
              |  |  |  +----------- Sinosolenaia_carinata **      
              |  |  |  |                                       
              |  |  |  |        +-- Sinosolenaia_oleivora **      
              |  |  +--+--------+                              
              |  |     |        +-- Sinosolenaia_recognita **     
              |  |     |                                       
              |  |     |  +-------- Gonidea_angulata **           
              |  |     +--+                                    
              |  |        |  +----- Microcondylaea_bonellii **    
              +--+        +--+                                 
                 |           |  +-- Leguminaia_wheatleyi **       
                 |           +--+                              
                 |              +-- Leguminaia **                 
                 |                                             
                 |        +-------- Pilsbryoconcha_exilis[3] **   
                 |  +-----+                                    
                 |  |     |  +----- Pilsbryoconcha_exilis[2] **   
                 |  |     +--+                                 
                 |  |        |  +-- Pilsbryoconcha_exilis[1] **   
                 |  |        +--+                              
                 |  |           +-- Pilsbryoconcha_carinifera **  
                 |  |                                          
                 +--+  +----------- Indopseudodon_indawgyiensis **
                    |  |                                       
                    |  |  +-------- Thaiconcha_callifera **       
                    |  +--+                                    
                    |  |  |  +----- Bineurus_exilis **            
                    |  |  +--+                                 
                    |  |     |  +-- Bineurus_mouhotii **          
                    +--+     +--+                              
                       |        +-- Bineurus_loeiensis **         
                       |                                       
                       |     +----- Pseudodon_vagulus **          
                       |  +--+                                 
                       |  |  |  +-- Pseudodon_mekongi **          
                       |  |  +--+                              
                       |  |     +-- Pseudodon_vondembuschianus ** 
                       +--+                                    
                          |     +-- Pseudodon_inoscularis **      
                          |  +--+                              
                          |  |  +-- Sundadontina_tumida **        
                          +--+                                 
                             |  +-- Nyeinchanconcha_nyeinchani ** 
                             +--+                              
                                +-- Sundadontina_cumingii **      

Campbell et al., 2005 [Publ 2660], fig. 1

iT tT bi riS oT roT roS iD
115 115 102 3 2 2 1 133.0
                                                     +-- Margaritifera_margaritifera   
  +--------------------------------------------------+                                 
  |                                                  +-- Cumberlandia_monodonta        
  |                                                                                    
  |  +-------------------------------------------------- Sinohyriopsis_cumingii **        
  |  |                                                                                 
  |  |                                      +----------- Anodonta_oregonensis **          
  |  |                                      |                                          
  |  +--------------------------------------+        +-- Anodonta_cygnea **               
  |  |                                      |  +-----+                                 
  |  |                                      |  |     +-- Alasmidonta_holstonia **         
--+  |                                      +--+                                       
  |  |                                         |  +----- Pyganodon_grandis **             
  |  |                                         +--+                                    
  |  |                                            |  +-- Lasmigona_costata **             
  |  |                                            +--+                                 
  |  |                                               +-- Pseudodontoideus_subvexus **     
  |  |                                                                                 
  |  |                                            +----- Gonidea_angulata **              
  |  |  +-----------------------------------------+                                    
  |  |  |                                         |  +-- Potomida_littoralis **           
  +--+  |                                         +--+                                 
     |  |                                            +-- Pronodularia_japanensis **       
     |  |                                                                              
     |  |                                            +-- Megalonaias_nervosa **           
     |  |                                +-----------+                                 
     |  |                                |           +-- Uniomerus_declivis **            
     |  |                                |                                             
     |  |  +-----------------------------+  +----------- Theliderma_metanevra **          
     |  |  |                             |  |                                          
     |  |  |                             |  |     +----- Quadrula_verrucosa **            
     |  |  |                             +--+  +--+                                    
     |  |  |                                |  |  |  +-- Quadrula_quadrula **             
     |  |  |                                |  |  +--+                                 
     |  |  |                                |  |     +-- Quadrula_nobilis **              
     |  |  |                                +--+                                       
     |  |  |                                   |  +----- Pustulosa_kieneriana **          
     |  |  |                                   |  |                                    
     |  |  |                                   |  |  +-- Cyclonaias_tuberculata **        
     +--+  |                                   +--+--+                                 
        |  |                                      |  +-- Pustulosa_succissa **            
        |  |                                      |                                    
        |  |                                      |  +-- Pustulosa_infucata **            
        |  |                                      +--+                                 
        |  |                                         +-- Pustulosa_kleiniana **           
        |  |                                                                           
        |  |        +----------------------------------- Elliptoideus_sloatianus **       
        |  |        |                                                                  
        |  |        |  +-------------------------------- Hemistena_lata **                
        |  |        |  |                                                               
        |  |  +-----+  |                          +----- Plethobasus_cyphyus **           
        |  |  |     |  |  +-----------------------+                                    
        |  |  |     |  |  |                       |  +-- Elliptio_arca **                 
        |  |  |     |  |  |                       +--+                                 
        |  |  |     |  |  |                          +-- Elliptio_crassidens **           
        |  |  |     +--+  |                                                            
        |  |  |        |  |                          +-- Eurynia_dilatata **              
        |  |  |        |  |  +-----------------------+                                 
        |  |  |        |  |  |                       +-- Parvaspina_collina **            
        +--+  |        |  |  |                                                         
           |  |        |  |  |                    +----- Pleuronaia_gibber **             
           |  |        |  |  +--------------------+                                    
           |  |        |  |  |                    |  +-- Pleuronaia_barnesiana **         
           |  |        +--+  |                    +--+                                 
           |  |           |  |                       +-- Pleuronaia_dolabelloides **      
           |  |           |  |                                                         
           |  |           |  |              +----------- Fusconaia_subrotunda **          
           |  |           |  |              |                                          
           |  |           |  |              |        +-- Fusconaia_cor **                 
           |  |           |  +--------------+  +-----+                                 
           |  |           |  |              |  |     +-- Fusconaia_cuneolus **            
           |  |           |  |              |  |                                       
           |  |           |  |              |  |     +-- Fusconaia_cerina[1] **           
           |  |           |  |              +--+     |                                 
           |  |           |  |                 |  +--+-- Fusconaia_cerina[2] **           
           |  |           +--+                 |  |  |                                 
           |  |              |                 |  |  +-- Fusconaia_flava **               
           |  |              |                 +--+                                    
           |  |              |                    |  +-- Fusconaia_burkei[1] **           
           |  |              |                    |  |                                 
           |  |              |                    |  +-- Fusconaia_burkei[2] **           
           |  |              |                    +--+                                 
           |  |              |                       +-- Fusconaia_burkei[3] **           
           |  |              |                       |                                 
           |  |              |                       +-- Fusconaia_escambia **            
           |  |              |                                                         
           |  |              |                       +-- Pleurobema_rubrum[2] **          
           |  |              |                    +--+                                 
           |  |              |                    |  +-- Pleurobema_cordatum **           
           |  |              |  +-----------------+                                    
           |  |              |  |                 |  +-- Pleurobema_rubrum[1] **          
           +--+              |  |                 +--+                                 
              |              |  |                    +-- Pleurobema_sintoxia **           
              |              +--+                                                      
              |                 |  +-------------------- Pleurobema_pyriforme **          
              |                 |  |                                                   
              |                 |  |                 +-- Pleurobema_georgianum[2] **      
              |                 |  |                 |                                 
              |                 +--+  +--------------+-- Pleurobema_georgianum[3] **      
              |                    |  |              |                                 
              |                    |  |              +-- Pleurobema_athearni **           
              |                    |  |                                                
              |                    |  |           +----- Pleurobema_decisum **            
              |                    +--+           |                                    
              |                       |  +--------+  +-- Pleurobema_hanleyianum[1] **     
              |                       |  |        |  |                                 
              |                       |  |        +--+-- Pleurobema_hanleyianum[2] **     
              |                       |  |           |                                 
              |                       |  |           +-- Pleurobema_georgianum[1] **      
              |                       +--+                                             
              |                          |           +-- Pleurobema_oviforme[1] **        
              |                          |           |                                 
              |                          |  +--------+-- Pleurobema_oviforme[2] **        
              |                          |  |        |                                 
              |                          +--+        +-- Pleurobema_clava **              
              |                             |                                          
              |                             |  +-------- Pleurobema_strodeanum **         
              |                             +--+                                       
              |                                |  +----- Pleurobema_rubellum **           
              |                                +--+                                    
              |                                   |  +-- Pleurobema_perovatum **          
              |                                   +--+                                 
              |                                      +-- Pleurobema_taitianum **          
              |                                                                        
              |                                   +----- Plectomerus_dombeyanus **        
              |  +--------------------------------+                                    
              |  |                                |  +-- Reginaia_ebenus **               
              |  |                                +--+                                 
              |  |                                   +-- Reginaia_rotulata **             
              |  |                                                                     
              |  |                                +----- Popenaias_popeii **              
              |  |  +-----------------------------+                                    
              |  |  |                             |  +-- Amblema_elliottii **             
              +--+  |                             +--+                                 
                 |  |                                +-- Amblema_plicata **               
                 |  |                                                                  
                 |  |                                +-- Toxolasma_parvum **              
                 |  |                          +-----+                                 
                 |  |                          |     +-- Toxolasma_texasiense **          
                 |  |  +-----------------------+                                       
                 |  |  |                       |  +----- Obliquaria_reflexa **            
                 +--+  |                       +--+                                    
                    |  |                          |  +-- Cyrtonaias_tampicoensis **       
                    |  |                          +--+                                 
                    |  |                             +-- Glebula_rotundata **             
                    |  |                                                               
                    |  |                             +-- Potamilus_fragilis **            
                    |  |                          +--+                                 
                    |  |                          |  +-- Potamilus_leptodon **            
                    |  |              +-----------+                                    
                    |  |              |           |  +-- Potamilus_alatus **              
                    |  |              |           +--+                                 
                    +--+              |              +-- Potamilus_purpuratus **          
                       |              |                                                
                       |  +-----------+              +-- Sagittunio_nasutus **            
                       |  |           |           +--+                                 
                       |  |           |           |  +-- Medionidus_conradicus **         
                       |  |           |  +--------+                                    
                       |  |           |  |        |  +-- Ptychobranchus_fasciolaris **    
                       |  |           |  |        +--+                                 
                       |  |           +--+           +-- Venustaconcha_ellipsiformis[2] **
                       |  |              |                                             
                       |  |              |  +----------- Medionidus_acutissimus **        
                       |  |              +--+                                          
                       |  |                 |  +-------- Lemiox_rimosus **                
                       |  |                 +--+                                       
                       +--+                    |  +----- Dromus_dromas **                 
                          |                    +--+                                    
                          |                       |  +-- Cyprogenia_aberti **             
                          |                       +--+                                 
                          |                          +-- Cyprogenia_stegaria **           
                          |                                                            
                          |                          +-- Ellipsaria_lineolata **          
                          |  +-----------------------+                                 
                          |  |                       +-- Truncilla_truncata **            
                          |  |                                                         
                          |  |  +----------------------- Villosa_villosa **               
                          |  |  |                                                      
                          |  |  |                    +-- Obovaria_olivaria **             
                          +--+  |     +--------------+                                 
                             |  |     |              +-- Venustaconcha_ellipsiformis[1] **
                             |  |     |                                                
                             |  |  +--+           +----- Obovaria_unicolor **             
                             |  |  |  |  +--------+                                    
                             |  |  |  |  |        |  +-- Obovaria_arkansasensis **        
                             +--+  |  |  |        +--+                                 
                                |  |  +--+           +-- Obovaria_subrotunda **           
                                |  |     |                                             
                                |  |     |  +----------- Epioblasma_brevidens **          
                                |  |     +--+                                          
                                |  |        |  +-------- Epioblasma_triquetra **          
                                |  |        +--+                                       
                                |  |           |  +----- Epioblasma_walkeri[2] **         
                                |  |           +--+                                    
                                |  |              |  +-- Epioblasma_capsaeformis **       
                                +--+              +--+                                 
                                   |                 +-- Epioblasma_walkeri[1] **         
                                   |                                                   
                                   |                 +-- Ligumia_recta **                 
                                   |        +--------+                                 
                                   |        |        +-- Cambarunio_iris **               
                                   |     +--+                                          
                                   |     |  |  +-------- Lampsilis_teres **               
                                   |     |  +--+                                       
                                   |     |     |  +----- Ortmanniana_ligamentina **       
                                   |     |     +--+                                    
                                   |     |        |  +-- Lampsilis_siliquoidea **         
                                   |     |        +--+                                 
                                   |     |           +-- Leaunio_vanuxemensis **          
                                   +-----+                                             
                                         |           +-- Lampsilis_cardium **             
                                         |        +--+                                 
                                         |        |  +-- Lampsilis_ovata **               
                                         |  +-----+                                    
                                         |  |     |  +-- Ortmanniana_pectorosa **         
                                         |  |     +--+                                 
                                         |  |        +-- Lampsilis_ornata **              
                                         |  |                                          
                                         +--+     +----- Hamiota_australis[2] **          
                                            |  +--+                                    
                                            |  |  |  +-- Hamiota_australis[1] **          
                                            |  |  +--+                                 
                                            |  |     +-- Hamiota_subangulata **           
                                            +--+                                       
                                               |  +----- Hamiota_altilis[2] **            
                                               |  |                                    
                                               |  |  +-- Hamiota_altilis[1] **            
                                               +--+--+                                 
                                                  |  +-- Hamiota_perovalis[1] **          
                                                  |                                    
                                                  |  +-- Hamiota_altilis[3] **            
                                                  +--+                                 
                                                     +-- Hamiota_perovalis[2] **          

Bolotov et al., 2020 [Publ 4100], fig. 1

iT tT bi riS oT roT roS iD
115 115 98 2 5 5 1 152.0
                                            +-- Gibbosula_laosensis          
                                            |                                
                                            +-- Margaritifera_dahurica       
                                            |                                
  +-----------------------------------------+-- Margaritifera_margaritifera  
  |                                         |                                
  |                                         +-- Margaritifera_laevis         
  |                                         |                                
  |                                         +-- Margaritifera_middendorffi   
  |                                                                          
  |                                   +-------- Trapezidens_exolescens **       
  |                                +--+                                      
  |                                |  |  +----- Trapezidens_scutum **           
  |                                |  +--+                                   
  |                                |     |  +-- Trapezidens_dolichorhynchus **  
  |     +--------------------------+     +--+                                
  |     |                          |        +-- Trapezidens_angustior **        
  |     |                          |                                         
  |     |                          |  +-------- Lamellidens_savadiensis **      
  |     |                          +--+                                      
  |     |                             |  +----- Lamellidens_generosus **        
  |     |                             +--+                                   
  |     |                                |  +-- Lamellidens_ferrugineus **      
  |     |                                +--+                                
  |     |                                   +-- Lamellidens_mainwaringi **      
  |     |                                                                    
  |     |                          +----------- Parreysia_keralaensis **        
  |     |                       +--+                                         
  |  +--+                       |  |  +-------- Parreysia_plagiosoma **         
  |  |  |                       |  +--+                                      
  |  |  |                       |     |  +----- Parreysia_nagpoorensis[2] **    
  |  |  |                       |     +--+                                   
  |  |  |                       |        |  +-- Parreysia_nagpoorensis[1] **    
  |  |  |  +--------------------+        +--+                                
  |  |  |  |                    |           +-- Parreysia_rakhinensis **        
  |  |  |  |                    |                                            
  |  |  |  |                    |           +-- Leoparreysia_burmana **         
  |  |  |  |                    |     +-----+                                
  |  |  |  |                    |     |     +-- Leoparreysia_whitteni **        
  |  |  |  |                    +-----+                                      
  |  |  |  |                          |  +----- Leoparreysia_tavoyensis **      
  |  |  |  |                          +--+                                   
  |  |  +--+                             |  +-- Leoparreysia_olivacea **        
  |  |     |                             +--+                                
  |  |     |                                +-- Leoparreysia_canefrii **        
  |  |     |                                                                 
  |  |     |                                +-- Coelatura_aegyptiaca **         
--+  |     |  +-----------------------------+                                
  |  |     |  |                             +-- Nitia_teretiuscula **           
  |  |     |  |                                                              
  |  |     |  |                    +----------- Indonaia_shurtleffiana **       
  |  |     |  |  +-----------------+                                         
  |  |     +--+  |                 |  +-------- Indonaia_rugosa **              
  |  |        |  |                 +--+                                      
  |  |        |  |                    |  +----- Indonaia_andersoniana **        
  |  |        |  |                    +--+                                   
  |  |        |  |                       |  +-- Indonaia_subclathrata **        
  |  |        +--+                       +--+                                
  |  |           |                          +-- Indonaia_caerulea **            
  |  |           |                                                           
  |  |           |  +-------------------------- Indochinella_pugio **           
  |  |           |  |                                                        
  |  |           |  |                 +-------- Radiatula_crispisulcata **      
  |  |           |  |  +--------------+                                      
  |  |           +--+  |              |  +----- Radiatula_mouhoti **            
  |  |              |  |              +--+                                   
  |  |              |  |                 |  +-- Radiatula_chaudhurii **         
  |  |              |  |                 +--+                                
  |  |              +--+                    +-- Radiatula_myitkyinae **         
  |  |                 |                                                     
  |  |                 |  +-------------------- Scabiellus_songkramensis **     
  |  |                 |  |                                                  
  |  |                 |  |                 +-- Unionetta_fabagina **           
  |  |                 +--+  +--------------+                                
  |  |                    |  |              +-- Harmandia_somboriensis **       
  |  |                    |  |                                               
  |  |                    +--+  +-------------- Scabies_nucleus **              
  |  |                       |  |                                            
  |  |                       |  |           +-- Scabies_crispata **             
  |  |                       +--+  +--------+                                
  |  |                          |  |        +-- Scabies_mandarinus **           
  |  |                          |  |                                         
  |  |                          +--+        +-- Scabies_humilis **              
  |  |                             |  +-----+                                
  |  |                             |  |     +-- Scabies_pilata **               
  |  |                             +--+                                      
  |  |                                |  +----- Scabies_anceps **               
  |  |                                +--+                                   
  |  |                                   |  +-- Scabies_phaselus **             
  |  |                                   +--+                                
  +--+                                      +-- Scabies_scobinatus **           
     |                                                                       
     |                             +----------- Hyriopsis_desowitzi **          
     |                             |                                         
     |                          +--+  +-------- Rectidens_sumatrensis **        
     |                          |  |  |                                      
     |                          |  +--+     +-- Hyriopsis_khoratensis **        
     |                          |     |  +--+                                
     |                          |     |  |  +-- Hyriopsis_bialata **            
     |                          |     +--+                                   
     |                          |        |  +-- Hyriopsis_myersiana **          
     |                 +--------+        +--+                                
     |                 |        |           +-- Hyriopsis_panhai **             
     |                 |        |                                            
     |                 |        |           +-- Ensidens_telus **               
     |                 |        |     +-----+                                
     |                 |        |     |     +-- Ensidens_ingallsianus **        
     |                 |        |     |                                      
     |                 |        +-----+     +-- Ensidens_jaculus **             
     |                 |              |  +--+                                
     |                 |              |  |  +-- Ensidens_spiculus **            
     |                 |              +--+                                   
     |                 |                 |  +-- Ensidens_dugasti **             
     |                 |                 +--+                                
     |                 |                    +-- Ensidens_sagittarius **         
     |           +-----+                                                     
     |           |     |                    +-- Trapezoideus_lenya **           
     |           |     |              +-----+                                
     |           |     |              |     +-- Trapezoideus_foliaceus **       
     |           |     |  +-----------+                                      
     |           |     |  |           |  +----- Physunio_superbus **            
     |           |     |  |           +--+                                   
     |           |     |  |              |  +-- Physunio_modelli **             
     |           |     |  |              +--+                                
     |           |     |  |                 +-- Physunio **                     
     |           |     |  |                                                  
     |           |     |  |                 +-- Yaukthwa_elongatula **          
     |           |     |  |     +-----------+                                
     |           |     |  |     |           +-- Yaukthwa_baniensis **           
     |           |     +--+     |                                            
     |           |        |  +--+        +----- Yaukthwa_inlenensis **          
     |           |        |  |  |  +-----+                                   
     |           |        |  |  |  |     |  +-- Yaukthwa_avaensis **            
     |           |        |  |  |  |     +--+                                
     |           |        |  |  +--+        +-- Yaukthwa_paiensis **            
     |           |        |  |     |                                         
     |           |        |  |     |  +-------- Yaukthwa_panhai **              
     |           |        |  |     +--+                                      
     |           |        |  |        |  +----- Yaukthwa_rectangularis **       
     |           |        +--+        +--+                                   
     |           |           |           |  +-- Yaukthwa_zayleymanensis **      
     |           |           |           +--+                                
     |           |           |              +-- Yaukthwa_nesemanni **           
     |           |           |                                               
     |           |           |           +----- Lens_misellus **                
     |           |           |     +-----+                                   
     |           |           |     |     |  +-- Lens_contradens **              
     |           |           |     |     +--+                                
     |           |           |     |        +-- Lens_micropterus **             
     |           |           +-----+                                         
     |           |                 |     +----- Lens_pallegoixi **              
     |           |                 |  +--+                                   
     +-----------+                 |  |  |  +-- Lens_comptus **                 
                 |                 |  |  +--+                                
                 |                 +--+     +-- Lens_novoselovi **              
                 |                    |                                      
                 |                    |  +----- Lens_maenamensis **             
                 |                    +--+                                   
                 |                       |  +-- Lens_eximius **                 
                 |                       +--+                                
                 |                          +-- Lens_rolfbrandti **             
                 |                                                           
                 |                          +-- Sinosolenaia_iridinea **        
                 |                          |                                
                 |                          +-- Sinosolenaia_recognita[1] **    
                 |                          |                                
                 |                          +-- Sinosolenaia_carinata **        
                 |                          |                                
                 |                          +-- Leguminaia_wheatleyi **         
                 |                          |                                
                 |                          +-- Microcondylaea_bonellii **      
                 |                          |                                
                 |                          +-- Gonidea_angulata **             
                 |                          |                                
                 |                          +-- Ptychorhynchus_pfisteri **      
                 |                          |                                
                 |                          +-- Parvasolenaia_rivularis **      
                 |                          |                                
                 |  +-----------------------+-- Lamprotula_leaii **             
                 |  |                       |                                
                 |  |                       +-- Lamprotula **                   
                 |  |                       |                                
                 |  |                       +-- Lamprotula_caveata **           
                 |  |                       |                                
                 |  |                       +-- Potomida_littoralis **          
                 |  |                       |                                
                 |  |                       +-- Pronodularia_japanensis **      
                 |  |                       |                                
                 |  |                       +-- Sinohyriopsis_cumingii **       
                 |  |                       |                                
                 +--+                       +-- Chamberlainia_somsakpanhai **   
                    |                       |                                
                    |                       +-- Sinosolenaia_recognita[2] **    
                    |                       |                                
                    |                       +-- Sinosolenaia_oleivora **        
                    |                                                        
                    |                 +-------- Indopseudodon_kayinensis **     
                    |  +--------------+                                      
                    |  |              |  +----- Indopseudodon_indawgyiensis **  
                    |  |              +--+                                   
                    |  |                 |  +-- Indopseudodon_salwenianus **    
                    |  |                 +--+                                
                    |  |                    +-- Indopseudodon_bogani **         
                    +--+                                                     
                       |                 +----- Bineurus_loeiensis **           
                       |  +--------------+                                   
                       |  |              |  +-- Bineurus_exilis **              
                       |  |              +--+                                
                       |  |                 +-- Bineurus_mouhotii **            
                       |  |                                                  
                       +--+  +----------------- Nyeinchanconcha_nyeinchani **   
                          |  |                                               
                          |  |              +-- Thaiconcha_callifera **         
                          |  |        +-----+                                
                          |  |        |     +-- Thaiconcha_munelliptica **      
                          +--+  +-----+                                      
                             |  |     |  +----- Pilsbryoconcha_exilis[2] **     
                             |  |     +--+                                   
                             |  |        |  +-- Pilsbryoconcha_exilis[1] **     
                             |  |        +--+                                
                             |  |           +-- Pilsbryoconcha **               
                             +--+                                            
                                |     +-------- Pseudodon_lenyanensis **        
                                |  +--+                                      
                                |  |  |  +----- Pseudodon_vagulus **            
                                |  |  +--+                                   
                                |  |     |  +-- Pseudodon_vondembuschianus **   
                                |  |     +--+                                
                                +--+        +-- Pseudodon_mekongi **            
                                   |                                         
                                   |        +-- Sundadontina_taskaevi **        
                                   |  +-----+                                
                                   |  |     +-- Sundadontina_brandti **         
                                   +--+                                      
                                      |  +----- Sundadontina_tumida **          
                                      +--+                                   
                                         |  +-- Sundadontina_cumingii **        
                                         +--+                                
                                            +-- Sundadontina_tanintharyiensis **

Bolotov et al., 2020 [Publ 4100], fig. S1

iT tT bi riS oT roT roS iD
114 114 100 2 0 0 0 151.0
                                         +-------- Trapezidens_exolescens **       
                                      +--+                                      
                                      |  |  +----- Trapezidens_scutum **           
                                      |  +--+                                   
                                      |     |  +-- Trapezidens_dolichorhynchus **  
     +--------------------------------+     +--+                                
     |                                |        +-- Trapezidens_angustior **        
     |                                |                                         
     |                                |  +-------- Lamellidens_savadiensis **      
     |                                +--+                                      
     |                                   |  +----- Lamellidens_generosus **        
     |                                   +--+                                   
     |                                      |  +-- Lamellidens_ferrugineus **      
     |                                      +--+                                
     |                                         +-- Lamellidens_mainwaringi **      
     |                                                                          
     |                                +----------- Parreysia_keralaensis **        
     |                             +--+                                         
  +--+                             |  |  +-------- Parreysia_nagpoorensis[2] **    
  |  |                             |  +--+                                      
  |  |                             |     |  +----- Parreysia_nagpoorensis[1] **    
  |  |                             |     +--+                                   
  |  |                             |        |  +-- Parreysia_rakhinensis **        
  |  |  +--------------------------+        +--+                                
  |  |  |                          |           +-- Parreysia_plagiosoma **         
  |  |  |                          |                                            
  |  |  |                          |           +-- Leoparreysia_burmana **         
  |  |  |                          |     +-----+                                
  |  |  |                          |     |     +-- Leoparreysia_whitteni **        
  |  |  |                          +-----+                                      
  |  |  |                                |  +----- Leoparreysia_olivacea **        
  |  |  |                                +--+                                   
  |  +--+                                   |  +-- Leoparreysia_tavoyensis **      
  |     |                                   +--+                                
  |     |                                      +-- Leoparreysia_canefrii **        
  |     |                                                                       
  |     |                                      +-- Coelatura_aegyptiaca **         
  |     |  +-----------------------------------+                                
  |     |  |                                   +-- Nitia_teretiuscula **           
  |     |  |                                                                    
  |     |  |                             +-------- Indonaia_shurtleffiana **       
  |     +--+  +--------------------------+                                      
  |        |  |                          |  +----- Indonaia_andersoniana **        
  |        |  |                          +--+                                   
  |        |  |                             |  +-- Indonaia_caerulea **            
  |        |  |                             +--+                                
  |        +--+                                +-- Indonaia_subclathrata **        
  |           |                                                                 
  |           |  +-------------------------------- Indochinella_pugio **           
  |           |  |                                                              
  |           |  |                          +----- Radiatula_mouhoti **            
  |           +--+  +-----------------------+                                   
  |              |  |                       |  +-- Radiatula_myitkyinae **         
  |              |  |                       +--+                                
  |              +--+                          +-- Radiatula_chaudhurii **         
  |                 |                                                           
  |                 |  +-------------------------- Radiatula_crispisulcata **      
  |                 |  |                                                        
  |                 +--+  +----------------------- Scabiellus_songkramensis **     
  |                    |  |                                                     
  |                    +--+  +-------------------- Harmandia_somboriensis **       
  |                       |  |                                                  
  |                       +--+  +----------------- Unionetta_fabagina **           
  |                          |  |                                               
  |                          +--+  +-------------- Scabies_mandarinus **           
  |                             |  |                                            
  |                             +--+  +----------- Scabies_crispata **             
  |                                |  |                                         
  |                                +--+  +-------- Scabies_nucleus **              
  |                                   |  |                                      
  |                                   +--+     +-- Scabies_humilis **              
  |                                      |  +--+                                
  |                                      |  |  +-- Scabies_pilata **               
  |                                      +--+                                   
  |                                         |  +-- Scabies_phaselus **             
  |                                         |  |                                
  |                                         +--+-- Scabies_scobinatus **           
  |                                            |                                
--+                                            +-- Scabies_anceps **               
  |                                                                             
  |                                +-------------- Hyriopsis_desowitzi **          
  |                             +--+                                            
  |                             |  |  +----------- Rectidens_sumatrensis **        
  |                             |  +--+                                         
  |                             |     |  +-------- Hyriopsis_panhai **             
  |                             |     +--+                                      
  |                             |        |  +----- Hyriopsis_myersiana **          
  |                             |        +--+                                   
  |                    +--------+           |  +-- Hyriopsis_khoratensis **        
  |                    |        |           +--+                                
  |                    |        |              +-- Hyriopsis_bialata **            
  |                    |        |                                               
  |                    |        |              +-- Ensidens_telus **               
  |                    |        |        +-----+                                
  |                    |        |        |     +-- Ensidens_ingallsianus **        
  |                    |        |        |                                      
  |                    |        +--------+     +-- Ensidens_dugasti **             
  |                    |                 |  +--+                                
  |                    |                 |  |  +-- Ensidens_sagittarius **         
  |                    |                 +--+                                   
  |                 +--+                    |  +-- Ensidens_spiculus **            
  |                 |  |                    +--+                                
  |                 |  |                       +-- Ensidens_jaculus **             
  |                 |  |                                                        
  |                 |  |                    +----- Physunio **                     
  |                 |  |  +-----------------+                                   
  |                 |  |  |                 |  +-- Physunio_modelli **             
  |                 |  |  |                 +--+                                
  |                 |  |  |                    +-- Physunio_superbus **            
  |                 |  |  |                                                     
  |                 |  |  |                    +-- Trapezoideus_lenya **           
  |                 |  |  |  +-----------------+                                
  |                 |  +--+  |                 +-- Trapezoideus_foliaceus **       
  |                 |     |  |                                                  
  |                 |     |  |              +----- Lens_misellus **                
  |                 |     |  |        +-----+                                   
  |                 |     |  |        |     |  +-- Lens_contradens **              
  |                 |     |  |        |     +--+                                
  |                 |     |  |        |        +-- Lens_micropterus **             
  |                 |     +--+  +-----+                                         
  |                 |        |  |     |     +----- Lens_maenamensis **             
  |                 |        |  |     |  +--+                                   
  |                 |        |  |     |  |  |  +-- Lens_eximius **                 
  |                 |        |  |     |  |  +--+                                
  |                 |        |  |     +--+     +-- Lens_rolfbrandti **             
  |                 |        |  |        |                                      
  |                 |        |  |        |  +----- Lens_pallegoixi **              
  |                 |        +--+        +--+                                   
  |                 |           |           |  +-- Lens_novoselovi **              
  |                 |           |           +--+                                
  |                 |           |              +-- Lens_comptus **                 
  |                 |           |                                               
  |                 |           |              +-- Yaukthwa_elongatula **          
  |                 |           |  +-----------+                                
  |                 |           |  |           +-- Yaukthwa_baniensis **           
  |                 |           |  |                                            
  |                 |           +--+        +----- Yaukthwa_inlenensis **          
  |                 |              |  +-----+                                   
  +-----------------+              |  |     |  +-- Yaukthwa_avaensis **            
                    |              |  |     +--+                                
                    |              +--+        +-- Yaukthwa_paiensis **            
                    |                 |                                         
                    |                 |  +-------- Yaukthwa_panhai **              
                    |                 +--+                                      
                    |                    |  +----- Yaukthwa_rectangularis **       
                    |                    +--+                                   
                    |                       |  +-- Yaukthwa_zayleymanensis **      
                    |                       +--+                                
                    |                          +-- Yaukthwa_nesemanni **           
                    |                                                           
                    |                          +-- Sinohyriopsis_cumingii **       
                    |                       +--+                                
                    |                       |  +-- Chamberlainia_somsakpanhai **   
                    |                       |                                   
                    |                    +--+  +-- Lamprotula_leaii **             
                    |                    |  |  |                                
                    |                    |  |  +-- Lamprotula **                   
                    |                    |  |  |                                
                    |                    |  +--+-- Lamprotula_caveata **           
                    |                    |     |                                
                    |                    |     +-- Potomida_littoralis **          
                    |                    |     |                                
                    |                    |     +-- Pronodularia_japanensis **      
                    |  +-----------------+                                      
                    |  |                 |     +-- Sinosolenaia_iridinea **        
                    |  |                 |     |                                
                    |  |                 |     +-- Sinosolenaia_recognita[1] **    
                    |  |                 |     |                                
                    |  |                 |     +-- Sinosolenaia_carinata **        
                    |  |                 |     |                                
                    |  |                 |     +-- Leguminaia_wheatleyi **         
                    |  |                 |     |                                
                    |  |                 |     +-- Microcondylaea_bonellii **      
                    |  |                 +-----+                                
                    |  |                       +-- Gonidea_angulata **             
                    |  |                       |                                
                    |  |                       +-- Ptychorhynchus_pfisteri **      
                    +--+                       |                                
                       |                       +-- Parvasolenaia_rivularis **      
                       |                       |                                
                       |                       +-- Sinosolenaia_recognita[2] **    
                       |                       |                                
                       |                       +-- Sinosolenaia_oleivora **        
                       |                                                        
                       |                 +-------- Indopseudodon_kayinensis **     
                       |  +--------------+                                      
                       |  |              |  +----- Indopseudodon_indawgyiensis **  
                       |  |              +--+                                   
                       |  |                 |  +-- Indopseudodon_salwenianus **    
                       |  |                 +--+                                
                       |  |                    +-- Indopseudodon_bogani **         
                       +--+                                                     
                          |                 +----- Bineurus_loeiensis **           
                          |  +--------------+                                   
                          |  |              |  +-- Bineurus_exilis **              
                          |  |              +--+                                
                          |  |                 +-- Bineurus_mouhotii **            
                          |  |                                                  
                          +--+  +----------------- Nyeinchanconcha_nyeinchani **   
                             |  |                                               
                             |  |              +-- Thaiconcha_callifera **         
                             |  |        +-----+                                
                             |  |        |     +-- Thaiconcha_munelliptica **      
                             +--+  +-----+                                      
                                |  |     |  +----- Pilsbryoconcha_exilis[2] **     
                                |  |     +--+                                   
                                |  |        |  +-- Pilsbryoconcha_exilis[1] **     
                                |  |        +--+                                
                                |  |           +-- Pilsbryoconcha **               
                                +--+                                            
                                   |     +-------- Pseudodon_lenyanensis **        
                                   |  +--+                                      
                                   |  |  |  +----- Pseudodon_vagulus **            
                                   |  |  +--+                                   
                                   |  |     |  +-- Pseudodon_vondembuschianus **   
                                   |  |     +--+                                
                                   +--+        +-- Pseudodon_mekongi **            
                                      |                                         
                                      |        +-- Sundadontina_tanintharyiensis **
                                      |  +-----+                                
                                      |  |     +-- Sundadontina_cumingii **        
                                      +--+                                      
                                         |  +----- Sundadontina_tumida **          
                                         +--+                                   
                                            |  +-- Sundadontina_taskaevi **        
                                            +--+                                
                                               +-- Sundadontina_brandti **         

Campbell et al., 2005 [Publ 2660], fig. 2

iT tT bi riS oT roT roS iD
113 113 91 3 2 2 1 129.0
                                               +-- Cumberlandia_monodonta        
  +--------------------------------------------+                                 
  |                                            +-- Margaritifera_margaritifera   
  |                                                                              
  |                                   +----------- Sinohyriopsis_cumingii **        
  |                                   |                                          
  |  +--------------------------------+  +-------- Pyganodon_grandis **             
  |  |                                |  |                                       
  |  |                                |  |     +-- Lasmigona_costata **             
  |  |                                +--+-----+                                 
  |  |                                   |     +-- Pseudodontoideus_subvexus **     
--+  |                                   |                                       
  |  |                                   |  +----- Alasmidonta_holstonia **         
  |  |                                   +--+                                    
  |  |                                      |  +-- Anodonta_cygnea **               
  |  |                                      +--+                                 
  |  |                                         +-- Anodonta_oregonensis **          
  |  |                                                                           
  |  |                                      +----- Gonidea_angulata **              
  |  |  +-----------------------------------+                                    
  |  |  |                                   |  +-- Potomida_littoralis **           
  |  |  |                                   +--+                                 
  +--+  |                                      +-- Pronodularia_japanensis **       
     |  |                                                                        
     |  |                    +-------------------- Megalonaias_nervosa **           
     |  |                    |                                                   
     |  |  +-----------------+  +----------------- Uniomerus_declivis **            
     |  |  |                 |  |                                                
     |  |  |                 +--+  +-------------- Theliderma_metanevra **          
     |  |  |                    |  |                                             
     |  |  |                    +--+  +----------- Quadrula_quadrula **             
     |  |  |                       |  |                                          
     |  |  |                       |  |        +-- Quadrula_nobilis **              
     |  |  |                       +--+  +-----+                                 
     |  |  |                          |  |     +-- Quadrula_verrucosa **            
     |  |  |                          |  |                                       
     |  |  |                          +--+  +----- Pustulosa_kieneriana **          
     |  |  |                             |  |                                    
     +--+  |                             |  |  +-- Cyclonaias_tuberculata **        
        |  |                             +--+--+                                 
        |  |                                |  +-- Pustulosa_succissa **            
        |  |                                |                                    
        |  |                                |  +-- Pustulosa_infucata **            
        |  |                                +--+                                 
        |  |                                   +-- Pustulosa_kleiniana **           
        |  |                                                                     
        |  |     +-------------------------------- Elliptoideus_sloatianus **       
        |  |     |                                                               
        |  |     |  +----------------------------- Plethobasus_cyphyus **           
        |  |     |  |                                                            
        |  |     |  +----------------------------- Parvaspina_collina **            
        |  |  +--+  |                                                            
        |  |  |  |  |                          +-- Elliptio_arca **                 
        |  |  |  |  +--------------------------+                                 
        |  |  |  |  |                          +-- Elliptio_crassidens **           
        |  |  |  |  |                                                            
        |  |  |  |  |  +-------------------------- Eurynia_dilatata **              
        |  |  |  +--+  |                                                         
        +--+  |     |  +-------------------------- Hemistena_lata **                
           |  |     |  |                                                         
           |  |     |  |                       +-- Pleurobema_rubrum[1] **          
           |  |     |  |                    +--+                                 
           |  |     |  |                    |  +-- Pleurobema_sintoxia **           
           |  |     |  |  +-----------------+                                    
           |  |     |  |  |                 |  +-- Pleurobema_rubrum[2] **          
           |  |     |  |  |                 +--+                                 
           |  |     +--+  |                    +-- Pleurobema_cordatum **           
           |  |        |  |                                                      
           |  |        |  |        +-------------- Fusconaia_subrotunda **          
           |  |        |  |        |                                             
           |  |        |  |        |           +-- Fusconaia_cor **                 
           |  |        |  +--------+  +--------+                                 
           |  |        |  |        |  |        +-- Fusconaia_cuneolus **            
           |  |        |  |        |  |                                          
           |  |        |  |        +--+        +-- Fusconaia_cerina **              
           |  |        |  |           |  +-----+                                 
           |  |        |  |           |  |     +-- Fusconaia_flava **               
           |  |        +--+           +--+                                       
           |  |           |              |  +----- Fusconaia_burkei[2] **           
           |  |           |              +--+                                    
           |  |           |                 |  +-- Fusconaia_escambia **            
           |  |           |                 +--+                                 
           |  |           |                    +-- Fusconaia_burkei[1] **           
           |  |           |                                                      
           |  |           |                 +----- Pleuronaia_gibber **             
           |  |           |  +--------------+                                    
           |  |           |  |              |  +-- Pleuronaia_barnesiana **         
           |  |           |  |              +--+                                 
           |  |           |  |                 +-- Pleuronaia_dolabelloides **      
           |  |           |  |                                                   
           |  |           |  |           +-------- Pleurobema_rubellum **           
           +--+           +--+  +--------+                                       
              |              |  |        |  +----- Pleurobema_strodeanum **         
              |              |  |        +--+                                    
              |              |  |           |  +-- Pleurobema_perovatum **          
              |              |  |           +--+                                 
              |              |  |              +-- Pleurobema_taitianum **          
              |              +--+                                                
              |                 |              +-- Pleurobema_oviforme[1] **        
              |                 |              |                                 
              |                 |  +-----------+-- Pleurobema_oviforme[2] **        
              |                 |  |           |                                 
              |                 |  |           +-- Pleurobema_clava **              
              |                 +--+                                             
              |                    |  +----------- Pleurobema_pyriforme **          
              |                    |  |                                          
              |                    +--+        +-- Pleurobema_athearni **           
              |                       |  +-----+                                 
              |                       |  |     +-- Pleurobema_georgianum[2] **      
              |                       +--+                                       
              |                          |  +----- Pleurobema_decisum **            
              |                          |  |                                    
              |                          +--+  +-- Pleurobema_hanleyianum[1] **     
              |                             |  |                                 
              |                             +--+-- Pleurobema_hanleyianum[2] **     
              |                                |                                 
              |                                +-- Pleurobema_georgianum[1] **      
              |                                                                  
              |                          +-------- Popenaias_popeii **              
              |                          |                                       
              |           +--------------+     +-- Amblema_elliottii **             
              |           |              |  +--+                                 
              |           |              |  |  +-- Amblema_plicata **               
              |           |              +--+                                    
              |           |                 |  +-- Reginaia_ebenus **               
              |           |                 |  |                                 
              |           |                 +--+-- Reginaia_rotulata[1] **          
              |           |                    |                                 
              |           |                    +-- Reginaia_rotulata[2] **          
              |           |                                                      
              |           |                 +----- Plectomerus_dombeyanus **        
              |           |  +--------------+                                    
              +-----------+  |              |  +-- Toxolasma_parvum **              
                          |  |              +--+                                 
                          |  |                 +-- Toxolasma_texasiense **          
                          |  |                                                   
                          |  |              +----- Obliquaria_reflexa **            
                          |  |  +-----------+                                    
                          |  |  |           |  +-- Cyrtonaias_tampicoensis **       
                          |  |  |           +--+                                 
                          |  |  |              +-- Glebula_rotundata **             
                          |  |  |                                                
                          |  |  |  +-------------- Sagittunio_nasutus **            
                          |  |  |  |                                             
                          +--+  |  +-------------- Leaunio_vanuxemensis **          
                             |  |  |                                             
                             |  |  |           +-- Ellipsaria_lineolata **          
                             |  |  +-----------+                                 
                             |  |  |           +-- Truncilla_truncata **            
                             |  |  |                                             
                             |  |  |        +----- Lampsilis_teres **               
                             |  |  +--------+                                    
                             |  |  |        |  +-- Ortmanniana_ligamentina **       
                             |  |  |        +--+                                 
                             |  |  |           +-- Lampsilis_siliquoidea **         
                             |  |  |                                             
                             |  |  |        +----- Villosa_villosa **               
                             |  |  +--------+                                    
                             +--+  |        |  +-- Ligumia_recta **                 
                                |  |        +--+                                 
                                |  |           +-- Cambarunio_iris **               
                                |  |                                             
                                |  |     +-------- Ortmanniana_pectorosa **         
                                |  +-----+                                       
                                |  |     |  +----- Lampsilis_ornata **              
                                |  |     +--+                                    
                                |  |        |  +-- Lampsilis_cardium **             
                                |  |        +--+                                 
                                |  |           +-- Lampsilis_ovata **               
                                |  |                                             
                                |  |           +-- Potamilus_fragilis **            
                                |  |        +--+                                 
                                |  |        |  +-- Potamilus_leptodon **            
                                |  +--------+                                    
                                |  |        |  +-- Potamilus_alatus **              
                                |  |        +--+                                 
                                |  |           +-- Potamilus_purpuratus **          
                                |  |                                             
                                |  |           +-- Ptychobranchus_fasciolaris **    
                                +--+  +--------+                                 
                                   |  |        +-- Venustaconcha_ellipsiformis[1] **
                                   |  |                                          
                                   +--+  +-------- Medionidus_conradicus **         
                                   |  |  |                                       
                                   |  |  |     +-- Lemiox_rimosus **                
                                   |  +--+-----+                                 
                                   |     |     +-- Medionidus_acutissimus **        
                                   |     |                                       
                                   |     |  +----- Dromus_dromas **                 
                                   |     +--+                                    
                                   |        |  +-- Cyprogenia_aberti **             
                                   |        +--+                                 
                                   |           +-- Cyprogenia_stegaria **           
                                   |                                             
                                   |        +----- Hamiota_australis[2] **          
                                   |  +-----+                                    
                                   |  |     |  +-- Hamiota_australis[1] **          
                                   |  |     +--+                                 
                                   |  |        +-- Hamiota_subangulata **           
                                   +--+                                          
                                   |  |        +-- Hamiota_altilis[3] **            
                                   |  |  +-----+                                 
                                   |  |  |     +-- Hamiota_perovalis[2] **          
                                   |  +--+                                       
                                   |     |  +----- Hamiota_altilis[2] **            
                                   |     +--+                                    
                                   |        |  +-- Hamiota_altilis[1] **            
                                   |        +--+                                 
                                   |           +-- Hamiota_perovalis[1] **          
                                   |                                             
                                   |  +----------- Obovaria_olivaria **             
                                   |  |                                          
                                   |  +----------- Venustaconcha_ellipsiformis[2] **
                                   |  |                                          
                                   |  |     +----- Obovaria_unicolor **             
                                   |  +-----+                                    
                                   +--+     |  +-- Obovaria_arkansasensis **        
                                      |     +--+                                 
                                      |        +-- Obovaria_subrotunda **           
                                      |                                          
                                      |        +-- Epioblasma_brevidens **          
                                      |  +-----+                                 
                                      |  |     +-- Epioblasma_triquetra **          
                                      +--+                                       
                                         |  +----- Epioblasma_walkeri[1] **         
                                         +--+                                    
                                            |  +-- Epioblasma_walkeri[2] **         
                                            +--+                                 
                                               +-- Epioblasma_capsaeformis **       

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
111 111 91 4 11 2 1 138.0
                                      +-- Coelatura_aegyptiaca **       
                                      |                              
  +-----------------------------------+-- Leoparreysia_tavoyensis **    
  |                                   |                              
  |                                   +-- Radiatula_mouhoti **          
  |                                                                  
  |                                   +-- Lens_contradens **            
  |                                   |                              
  |                                   +-- Ensidens_sagittarius **       
  |                                   |                              
  |           +-----------------------+-- Physunio_modelli **           
  |           |                       |                              
  |           |                       +-- Rectidens_sumatrensis **      
  |           |                       |                              
  |           |                       +-- Yaukthwa_nesemanni **         
  |           |                                                      
  |           |                       +-- Amblema_plicata **            
  |           |                       |                              
  |           |                       +-- Elliptio_complanata **        
  |           |  +--------------------+                              
  |  +--------+  |                    +-- Lampsilis_cardium **          
  |  |        |  |                    |                              
  |  |        |  |                    +-- Quadrula_quadrula **          
  |  |        |  |                                                   
  |  |        |  |                 +----- Chamberlainia_somsakpanhai ** 
  |  |        |  |     +-----------+                                 
  |  |        |  |     |           |  +-- Sinohyriopsis_cumingii **     
  |  |        |  |     |           +--+                              
  |  |        |  |  +--+              +-- Sinohyriopsis_schlegelii **   
  |  |        |  |  |  |                                             
  |  |        |  |  |  |  +-------------- Pronodularia_japanensis **    
  |  |        +--+  |  |  |                                          
  |  |           |  |  +--+  +----------- Potomida_littoralis **        
  |  |           |  |     |  |                                       
  |  |           |  |     +--+  +-------- Lamprotula_caveata **         
  |  |           |  |        |  |                                    
--+  |           |  |        +--+  +----- Lamprotula_leaii[2] **        
  |  |           |  |           |  |                                 
  |  |           |  |           +--+----- Lamprotula_leaii[3] **        
  |  |           |  |              |                                 
  |  |           |  |              |  +-- Lamprotula_seomjinensis **    
  |  |           |  |              +--+                              
  |  |           |  |                 +-- Lamprotula_leaii[1] **        
  |  |           +--+                                                
  |  |              |                 +-- Indopseudodon_indawgyiensis **
  |  |              |        +--------+                              
  |  |              |        |        +-- Indopseudodon_salwenianus **  
  |  |              |        |                                       
  |  |              |  +-----+        +-- Pilsbryoconcha_exilis **      
  |  |              |  |     |  +-----+                              
  |  |              |  |     |  |     +-- Pilsbryoconcha_carinifera **  
  |  |              |  |     +--+                                    
  |  |              |  |        |  +----- Pseudodon_mekongi **          
  |  |              |  |        |  |                                 
  |  |              |  |        +--+  +-- Nyeinchanconcha_nyeinchani ** 
  |  |              |  |           |  |                              
  |  |              |  |           +--+-- Bineurus_mouhotii **          
  |  |              |  |              |                              
  |  |              +--+              +-- Sundadontina_tumida **        
  |  |                 |                                             
  |  |                 |        +-------- Inversidens_brandtii **       
  |  |                 |        |                                    
  |  |                 |  +-----+     +-- Obovalis_omiensis **          
  |  |                 |  |     |  +--+                              
  |  |                 |  |     |  |  +-- Ptychorhynchus_pfisteri **    
  |  |                 |  |     +--+                                 
  |  |                 |  |        |  +-- Parvasolenaia_rivularis **    
  |  |                 |  |        +--+                              
  |  |                 +--+           +-- Koreosolenaia_sitgyensis **   
  |  |                    |                                          
  +--+                    |        +----- Gonidea_angulata **           
     |                    |  +-----+                                 
     |                    |  |     |  +-- Leguminaia_wheatleyi **       
     |                    |  |     +--+                              
     |                    +--+        +-- Microcondylaea_bonellii **    
     |                       |                                       
     |                       |  +-------- Sinosolenaia_carinata **      
     |                       +--+                                    
     |                          |  +----- Sinosolenaia_recognita **     
     |                          +--+                                 
     |                             |  +-- Sinosolenaia_iridinea **      
     |                             +--+                              
     |                                +-- Sinosolenaia_oleivora **      
     |                                                               
     |  +-------------------------------- Acuticosta_chinensis **       
     |  |                                                            
     |  +-------------------------------- Lepidodesma_languilati **     
     |  |                                                            
     |  |                    +----------- Aculamprotula_tortuosa **     
     |  |     +--------------+                                       
     |  |     |              |  +-------- Aculamprotula_scripta **      
     |  |     |              +--+                                    
     |  |     |                 |  +----- Aculamprotula_coreana **      
     |  |     |                 +--+                                 
     |  |     |                    |  +-- Aculamprotula_fibrosa **      
     |  |     |                    +--+                              
     |  +-----+                       +-- Aculamprotula_tientsinensis **
     |  |     |                                                      
     |  |     |  +----------------------- Middendorffinaia_mongolica ** 
     |  |     |  |                                                   
     |  |     |  |                 +----- Unio_gibbus **                
     |  |     |  |                 |                                 
     |  |     |  |  +--------------+----- Unio_tumidus **               
     |  |     +--+  |              |                                 
     |  |        |  |              |  +-- Unio_crassus **               
     |  |        |  |              +--+                              
     |  |        |  |                 +-- Unio_pictorum **              
     |  |        |  |                                                
     |  |        |  |     +-------------- Nodularia_breviconcha **      
     |  |        +--+  +--+                                          
     |  |           |  |  |  +----------- Nodularia[2] **               
     |  |           |  |  +--+                                       
     |  |           |  |     |  +-------- Nodularia_nuxpersicae **      
     |  |           |  |     +--+                                    
     |  |           |  |        |  +----- Nodularia_nipponensis **      
     |  |           |  |        +--+                                 
     |  |           |  |           |  +-- Nodularia_douglasiae **       
     |  |           +--+           +--+                              
     |  |              |              +-- Nodularia[1] **               
     |  |              |                                             
     +--+              |              +-- Schistodesmus_lampreyanus **  
        |              |  +-----------+                              
        |              |  |           +-- Schistodesmus_spinosus **     
        |              |  |                                          
        |              |  |        +----- Cuneopsis_rufescens **        
        |              |  +--------+                                 
        |              +--+        |  +-- Cuneopsis_heudei **           
        |                 |        +--+                              
        |                 |           +-- Cuneopsis_celtiformis **      
        |                 |                                          
        |                 |  +----------- Tchangsinaia_piscicula **     
        |                 +--+                                       
        |                    |  +-------- Inversiunio_jokohamensis **   
        |                    +--+                                    
        |                       |  +----- Inversiunio_verrucosus **     
        |                       +--+                                 
        |                          |  +-- Inversiunio_reinianus **      
        |                          +--+                              
        |                             +-- Inversiunio_yanagawensis **   
        |                                                            
        |              +----------------- Lanceolaria_oxyrhyncha **     
        |              |                                             
        |  +-----------+  +-------------- Lanceolaria_gladiola **       
        |  |           |  |                                          
        |  |           +--+           +-- Lanceolaria_bilirata **       
        |  |              |  +--------+                              
        |  |              |  |        +-- Lanceolaria_grayii **         
        |  |              +--+                                       
        |  |                 |  +-------- Lanceolaria_acrorrhyncha **   
        |  |                 +--+                                    
        |  |                    |  +----- Lanceolaria_kihirai **        
        |  |                    +--+                                 
        |  |                       |  +-- Lanceolaria_lanceolata **     
        |  |                       +--+                              
        |  |                          +-- Lanceolaria_triformis **      
        |  |                                                         
        |  |                    +-------- Pseudanodonta_complanata **   
        +--+                    |                                    
           |                    |  +----- Anodonta_cygnea **            
           |                    +--+                                 
           |                    |  |  +-- Anodonta_anatina **           
           |  +-----------------+  +--+                              
           |  |                 |     +-- Anodonta_nuttalliana **       
           |  |                 |                                    
           |  |                 |  +----- Pyganodon_grandis **          
           |  |                 |  |                                 
           |  |                 |  +----- Simpsonaias_ambigua **        
           |  |                 +--+                                 
           |  |                    +----- Strophitus_undulatus **       
           |  |                    |                                 
           |  |                    |  +-- Alasmidonta_marginata **      
           |  |                    +--+                              
           |  |                       +-- Lasmigona_costata **          
           +--+                                                      
              |                       +-- Cristaria_plicata **          
              |  +--------------------+                              
              |  |                    +-- Cristaria_clessini **         
              |  |                                                   
              |  |        +-------------- Anemina_arcaeformis **        
              |  |        |                                          
              |  +--------+  +----------- Amuranodonta_kijaensis **     
              |  |        |  |                                       
              |  |        +--+  +-------- Buldowskia_suifunica **       
              |  |           |  |                                    
              |  |           +--+     +-- Buldowskia_flavotincta **     
              +--+              |  +--+                              
                 |              |  |  +-- Buldowskia_shadini **         
                 |              +--+                                 
                 |                 |  +-- Buldowskia_iwakawai **        
                 |                 +--+                              
                 |                    +-- Buldowskia_kamiyai **         
                 |                                                   
                 |  +-------------------- Pletholophus_tenuis **        
                 |  |                                                
                 |  |           +-------- Beringiana_japonica **        
                 |  |  +--------+                                    
                 +--+  |        |  +----- Beringiana_beringiana **      
                    |  |        +--+                                 
                    |  |           |  +-- Beringiana_fukuharai **       
                    |  |           +--+                              
                    +--+              +-- Beringiana_gosannensis **     
                       |                                             
                       |              +-- Sinanodonta_angula **         
                       |  +-----------+                              
                       |  |           +-- Sinanodonta_lucida **         
                       |  |                                          
                       +--+  +----------- Sinanodonta_schrenkii **      
                          |  |                                       
                          |  |        +-- Sinanodonta_calipygos **      
                          |  +--------+                              
                          +--+        +-- Sinanodonta_tumens **         
                             |                                       
                             |  +-------- Sinanodonta_woodiana[2] **    
                             |  |                                    
                             +--+  +----- Sinanodonta_elliptica **      
                                |  |                                 
                                +--+----- Sinanodonta_jourdyi **        
                                   |                                 
                                   |  +-- Sinanodonta_lauta **          
                                   +--+                              
                                      +-- Sinanodonta_woodiana[1] **    

Wu et al., 2023 [Publ 4339], fig. 4, S5-nt

iT tT bi riS oT roT roS iD
86 86 81 4 11 7 1 121.0
                                            +-- Gibbosula_crassa            
                                +-----------+                               
                                |           +-- Gibbosula_rochechouartii    
  +-----------------------------+                                           
  |                             |  +----------- Cumberlandia_monodonta      
  |                             +--+                                        
  |                                |  +-------- Pseudunio_marocanus         
  |                                +--+                                     
  |                                   |  +----- Margaritifera_falcata       
  |                                   +--+                                  
  |                                      |  +-- Margaritifera_dahurica      
  |                                      +--+                               
  |                                         +-- Margaritifera_margaritifera 
  |                                                                         
  |                                         +-- Quadrula_quadrula **           
  |              +--------------------------+                               
  |              |                          +-- Uniomerus_tetralasmus **       
  |              |                                                          
  |  +-----------+                          +-- Elliptio_complanata **         
  |  |           |  +-----------------------+                               
  |  |           |  |                       +-- Pleurobema_oviforme **         
  |  |           +--+                                                       
  |  |              |  +----------------------- Amblema_plicata **             
  |  |              |  |                                                    
  |  |              +--+  +-------------------- Popenaias_popeii **            
--+  |                 |  |                                                 
  |  |                 +--+  +----------------- Toxolasma_parvum **            
  |  |                    |  |                                              
  |  |                    |  |           +----- Potamilus_leptodon **          
  |  |                    +--+  +--------+                                  
  |  |                       |  |        |  +-- Potamilus_alatus **            
  |  |                       |  |        +--+                               
  |  |                       |  |           +-- Potamilus_streckersoni **      
  |  |                       +--+                                           
  |  |                          |  +----------- Leaunio_lienosus **            
  |  |                          |  |                                        
  |  |                          |  |     +----- Lampsilis_ornata **            
  |  |                          +--+  +--+                                  
  |  |                             |  |  |  +-- Lampsilis_cardium **           
  |  |                             |  |  +--+                               
  |  |                             +--+     +-- Lampsilis_cariosa **           
  |  |                                |                                     
  |  |                                |  +----- Venustaconcha_ellipsiformis ** 
  |  |                                +--+                                  
  |  |                                   |  +-- Lampsilis_powellii **          
  |  |                                   +--+                               
  |  |                                      +-- Lampsilis_siliquoidea **       
  +--+                                                                      
     |                                   +----- Chamberlainia_somsakpanhai **  
     |                 +-----------------+                                  
     |                 |                 |  +-- Sinohyriopsis_cumingii **      
     |                 |                 +--+                               
     |                 |                    +-- Sinohyriopsis_schlegelii **    
     |                 |                                                    
     |                 |                    +-- Hyriopsis_bialata **           
     |                 |                 +--+                               
     |  +--------------+                 |  +-- Rectidens_sumatrensis **       
     |  |              |  +--------------+                                  
     |  |              |  |              |  +-- Lens_contradens **             
     |  |              |  |              +--+                               
     |  |              |  |                 +-- Physunio_superbus **           
     |  |              |  |                                                 
     |  |              |  |        +----------- Potomida_littoralis **         
     |  |              +--+  +-----+                                        
     |  |                 |  |     |  +-------- Pronodularia_japanensis **     
     |  |                 |  |     +--+                                     
     |  |                 |  |        |  +----- Lamprotula_leaii **            
     |  |                 |  |        +--+                                  
     |  |                 |  |           |  +-- Lamprotula_caveata **          
     |  |                 +--+           +--+                               
     |  |                    |              +-- Lamprotula_cornuumlunae **     
     |  |                    |                                              
     |  |                    |              +-- Pseudodon_vondembuschianus **  
     |  |                    |  +-----------+                               
     |  |                    |  |           +-- Pilsbryoconcha_exilis **       
     |  |                    |  |                                           
     |  |                    +--+           +-- Ptychorhynchus_pfisteri **     
     +--+                       |  +--------+                               
        |                       |  |        +-- Microcondylaea_bonellii **     
        |                       +--+                                        
        |                          |  +-------- Parvasolenaia_rivularis **     
        |                          +--+                                     
        |                             |  +----- Sinosolenaia_carinata **       
        |                             +--+                                  
        |                                |  +-- Sinosolenaia_iridinea **       
        |                                +--+                               
        |                                   +-- Sinosolenaia_oleivora **       
        |                                                                   
        |  +----------------------------------- Lamellidens_marginalis **      
        |  |                                                                
        |  |                                +-- Lepidodesma_aligera **         
        |  |  +-----------------------------+                               
        |  |  |                             +-- Lepidodesma_languilati **      
        |  |  |                                                             
        |  |  |                       +-------- Acuticosta_chinensis **        
        |  |  |        +--------------+                                     
        |  |  |        |              |  +----- Lanceolaria_gladiola **        
        |  |  |        |              +--+                                  
        +--+  |        |                 |  +-- Lanceolaria_grayii **          
           |  |        |                 +--+                               
           |  |        |                    +-- Lanceolaria_lanceolata **      
           |  |        |                                                    
           |  |        |        +-------------- Cristaria_plicata **           
           |  |  +-----+        |                                           
           |  |  |     |  +-----+           +-- Anemina_arcaeformis **         
           |  |  |     |  |     |  +--------+                               
           |  |  |     |  |     |  |        +-- Anemina_euscaphys **           
           |  |  |     |  |     +--+                                        
           +--+  |     |  |        |  +-------- Beringiana_fukuharai **        
              |  |     |  |        +--+                                     
              |  |     |  |           |  +----- Sinanodonta_lucida **          
              |  |     |  |           +--+                                  
              |  |     +--+              |  +-- Sinanodonta_tumens **          
              |  |        |              +--+                               
              |  |        |                 +-- Sinanodonta_woodiana **        
              |  |        |                                                 
              |  |        |  +----------------- Anodonta_nuttalliana **        
              |  |        |  |                                              
              |  |        |  +----------------- Pseudanodonta_complanata **    
              |  |        |  |                                              
              |  |        +--+  +-------------- Anodonta_cygnea **             
              |  |           |  |                                           
              |  |           |  |           +-- Anodonta_exulcerata **         
              |  |           |  +-----------+                               
              |  |           |  |           +-- Anodonta_anatina **            
              |  |           +--+                                           
              +--+              |        +----- Pyganodon_grandis **           
                 |              |  +-----+                                  
                 |              |  |     |  +-- Utterbackia_imbecillis **      
                 |              |  |     +--+                               
                 |              +--+        +-- Utterbackia_peninsularis **    
                 |                 |                                        
                 |                 |  +-------- Lasmigona_complanata **        
                 |                 +--+                                     
                 |                    |  +----- Prolasmidonta_heterodon **     
                 |                    +--+                                  
                 |                       |  +-- Alasmidonta_varicosa **        
                 |                       +--+                               
                 |                          +-- Platynaias_compressa **        
                 |                                                          
                 |                    +-------- Aculamprotula_tortuosa **      
                 |                    |                                     
                 |  +-----------------+     +-- Aculamprotula_coreana **       
                 |  |                 |  +--+                               
                 |  |                 |  |  +-- Aculamprotula_tientsinensis ** 
                 |  |                 +--+                                  
                 |  |                    |  +-- Aculamprotula_polysticta **    
                 |  |                    +--+                               
                 |  |                       +-- Aculamprotula_scripta **       
                 |  |                                                       
                 +--+              +----------- Unio_tumidus **                
                    |              |                                        
                    |  +-----------+  +-------- Unio_crassus **                
                    |  |           |  |                                     
                    |  |           +--+  +----- Unio_elongatulus **            
                    |  |              |  |                                  
                    |  |              +--+  +-- Unio_delphinus **              
                    |  |                 |  |                               
                    |  |                 +--+-- Unio_mancus **                 
                    +--+                    |                               
                       |                    +-- Unio_pictorum **               
                       |                                                    
                       |                    +-- Pseudocuneopsis_capitata **    
                       |  +-----------------+                               
                       |  |                 +-- Pseudocuneopsis_sichuanensis **
                       |  |                                                 
                       +--+                 +-- Nodularia_breviconcha **       
                          |  +--------------+                               
                          |  |              +-- Nodularia_nipponensis **       
                          |  |                                              
                          +--+           +----- Tchangsinaia_piscicula **      
                             |  +--------+                                  
                             |  |        |  +-- Schistodesmus_lampreyanus[2] **
                             |  |        +--+                               
                             +--+           +-- Schistodesmus_spinosus **      
                                |                                           
                                |  +----------- Schistodesmus_lampreyanus[1] **
                                +--+                                        
                                   |  +-------- Cuneopsis_rufescens **         
                                   +--+                                     
                                      |  +----- Cuneopsis_celtiformis **       
                                      +--+                                  
                                         |  +-- Cuneopsis_demangei **          
                                         +--+                               
                                            +-- Cuneopsis_heudei **            

Wu et al., 2024 [Publ 4350], fig. 8a, S1

iT tT bi riS oT roT roS iD
86 86 83 4 6 2 1 122.0
                                         +-- Margaritifera_dahurica      
  +--------------------------------------+                               
  |                                      +-- Margaritifera_margaritifera 
  |                                                                      
  |                                      +-- Quadrula_quadrula **           
  |           +--------------------------+                               
  |           |                          +-- Uniomerus_tetralasmus **       
  |           |                                                          
  |  +--------+                          +-- Elliptio_complanata **         
  |  |        |  +-----------------------+                               
  |  |        |  |                       +-- Pleurobema_oviforme **         
  |  |        +--+                                                       
  |  |           |  +----------------------- Amblema_plicata **             
  |  |           |  |                                                    
  |  |           +--+  +-------------------- Popenaias_popeii **            
  |  |              |  |                                                 
  |  |              +--+  +----------------- Toxolasma_parvum **            
  |  |                 |  |                                              
  |  |                 |  |           +----- Potamilus_leptodon **          
--+  |                 +--+  +--------+                                  
  |  |                    |  |        |  +-- Potamilus_alatus **            
  |  |                    |  |        +--+                               
  |  |                    |  |           +-- Potamilus_streckersoni **      
  |  |                    +--+                                           
  |  |                       |  +----------- Leaunio_lienosus **            
  |  |                       |  |                                        
  |  |                       |  |     +----- Lampsilis_ornata **            
  |  |                       +--+  +--+                                  
  |  |                          |  |  |  +-- Lampsilis_cardium **           
  |  |                          |  |  +--+                               
  |  |                          +--+     +-- Lampsilis_cariosa **           
  |  |                             |                                     
  |  |                             |  +----- Venustaconcha_ellipsiformis ** 
  |  |                             +--+                                  
  |  |                                |  +-- Lampsilis_powellii **          
  |  |                                +--+                               
  |  |                                   +-- Lampsilis_siliquoidea **       
  |  |                                                                   
  +--+                                +----- Chamberlainia_somsakpanhai **  
     |           +--------------------+                                  
     |           |                    |  +-- Sinohyriopsis_cumingii **      
     |           |                    +--+                               
     |           |                       +-- Sinohyriopsis_schlegelii **    
     |           |                                                       
     |           |                       +-- Hyriopsis_bialata **           
     |           |                    +--+                               
     |  +--------+                    |  +-- Rectidens_sumatrensis **       
     |  |        |  +-----------------+                                  
     |  |        |  |                 |  +-- Lens_contradens **             
     |  |        |  |                 +--+                               
     |  |        |  |                    +-- Physunio_superbus **           
     |  |        |  |                                                    
     |  |        |  |           +----------- Potomida_littoralis **         
     |  |        +--+  +--------+                                        
     |  |           |  |        |  +-------- Pronodularia_japanensis **     
     |  |           |  |        +--+                                     
     |  |           |  |           |  +----- Lamprotula_leaii **            
     |  |           |  |           +--+                                  
     |  |           |  |              |  +-- Lamprotula_caveata **          
     |  |           +--+              +--+                               
     |  |              |                 +-- Lamprotula_cornuumlunae **     
     |  |              |                                                 
     |  |              |                 +-- Pseudodon_vondembuschianus **  
     |  |              |  +--------------+                               
     |  |              |  |              +-- Pilsbryoconcha_exilis **       
     |  |              +--+                                              
     |  |                 |  +-------------- Inversidens_rentianensis **    
     |  |                 |  |                                           
     +--+                 +--+           +-- Parvasolenaia_rivularis **     
        |                    |  +--------+                               
        |                    |  |        +-- Ptychorhynchus_pfisteri **     
        |                    +--+                                        
        |                       |  +-------- Microcondylaea_bonellii **     
        |                       +--+                                     
        |                          |  +----- Sinosolenaia_carinata **       
        |                          +--+                                  
        |                             |  +-- Sinosolenaia_iridinea **       
        |                             +--+                               
        |                                +-- Sinosolenaia_oleivora **       
        |                                                                
        |  +-------------------------------- Lamellidens_marginalis **      
        |  |                                                             
        |  |                             +-- Lepidodesma_aligera **         
        |  |  +--------------------------+                               
        |  |  |                          +-- Lepidodesma_languilati **      
        |  |  |                                                          
        |  |  |                    +-------- Aculamprotula_tortuosa **      
        |  |  |                    |                                     
        |  |  |     +--------------+     +-- Aculamprotula_coreana **       
        |  |  |     |              |  +--+                               
        +--+  |     |              |  |  +-- Aculamprotula_tientsinensis ** 
           |  |     |              +--+                                  
           |  |     |                 |  +-- Aculamprotula_polysticta **    
           |  |     |                 +--+                               
           |  |     |                    +-- Aculamprotula_scripta **       
           |  |     |                                                    
           |  |  +--+        +-------------- Unio_tumidus **                
           |  |  |  |  +-----+                                           
           |  |  |  |  |     |  +----------- Unio_crassus **                
           |  |  |  |  |     +--+                                        
           +--+  |  |  |        |  +-------- Unio_elongatulus **            
              |  |  |  |        +--+                                     
              |  |  |  |           |  +----- Unio_pictorum **               
              |  |  |  |           +--+                                  
              |  |  |  |              |  +-- Unio_delphinus **              
              |  |  +--+              +--+                               
              |  |     |                 +-- Unio_mancus **                 
              |  |     |                                                 
              |  |     |                 +-- Nodularia_breviconcha **       
              |  |     |  +--------------+                               
              |  |     |  |              +-- Nodularia_douglasiae **        
              |  |     |  |                                              
              |  |     |  |              +-- Pseudocuneopsis_capitata **    
              |  |     |  +--------------+                               
              |  |     +--+              +-- Pseudocuneopsis_sichuanensis **
              |  |        |                                              
              |  |        |              +-- Schistodesmus_lampreyanus **   
              |  |        |  +-----------+                               
              +--+        |  |           +-- Schistodesmus_spinosus **      
                 |        +--+                                           
                 |           |  +----------- Tchangsinaia_piscicula **      
                 |           +--+                                        
                 |              |  +-------- Cuneopsis_rufescens **         
                 |              +--+                                     
                 |                 |  +----- Cuneopsis_celtiformis **       
                 |                 +--+                                  
                 |                    |  +-- Cuneopsis_demangei **          
                 |                    +--+                               
                 |                       +-- Cuneopsis_heudei **            
                 |                                                       
                 |                 +-------- Acuticosta_chinensis **        
                 |  +--------------+                                     
                 |  |              |  +----- Lanceolaria_gladiola **        
                 |  |              +--+                                  
                 |  |                 |  +-- Lanceolaria_grayii **          
                 |  |                 +--+                               
                 |  |                    +-- Lanceolaria_lanceolata **      
                 |  |                                                    
                 |  |        +-------------- Cristaria_plicata **           
                 |  |        |                                           
                 +--+  +-----+           +-- Anemina_arcaeformis **         
                    |  |     |  +--------+                               
                    |  |     |  |        +-- Anemina_euscaphys **           
                    |  |     +--+                                        
                    |  |        |  +-------- Beringiana_fukuharai **        
                    |  |        +--+                                     
                    |  |           |  +----- Sinanodonta_lucida **          
                    |  |           +--+                                  
                    |  |              |  +-- Sinanodonta_tumens **          
                    +--+              +--+                               
                       |                 +-- Sinanodonta_woodiana **        
                       |                                                 
                       |           +-------- Pseudanodonta_complanata **    
                       |  +--------+                                     
                       |  |        |  +----- Anodonta_anatina **            
                       |  |        +--+                                  
                       |  |           |  +-- Anodonta_cygnea **             
                       |  |           +--+                               
                       +--+              +-- Anodonta_exulcerata **         
                          |                                              
                          |  +-------------- Anodonta_nuttalliana **        
                          |  |                                           
                          |  |        +----- Pyganodon_grandis **           
                          +--+  +-----+                                  
                             |  |     |  +-- Utterbackia_imbecillis **      
                             |  |     +--+                               
                             +--+        +-- Utterbackia_peninsularis **    
                                |                                        
                                |  +-------- Lasmigona_complanata **        
                                +--+                                     
                                   |  +----- Prolasmidonta_heterodon **     
                                   +--+                                  
                                      |  +-- Alasmidonta_varicosa **        
                                      +--+                               
                                         +-- Platynaias_compressa **        

Wu et al., 2024 [Publ 4350], fig. 8b, S2

iT tT bi riS oT roT roS iD
86 86 82 4 6 2 1 122.0
                                         +-- Margaritifera_dahurica      
  +--------------------------------------+                               
  |                                      +-- Margaritifera_margaritifera 
  |                                                                      
  |                                      +-- Quadrula_quadrula **           
  |           +--------------------------+                               
  |           |                          +-- Uniomerus_tetralasmus **       
  |           |                                                          
  |  +--------+                          +-- Elliptio_complanata **         
  |  |        |  +-----------------------+                               
  |  |        |  |                       +-- Pleurobema_oviforme **         
  |  |        +--+                                                       
  |  |           |  +----------------------- Amblema_plicata **             
  |  |           |  |                                                    
  |  |           +--+  +-------------------- Popenaias_popeii **            
  |  |              |  |                                                 
  |  |              +--+  +----------------- Toxolasma_parvum **            
  |  |                 |  |                                              
  |  |                 |  |           +----- Potamilus_leptodon **          
--+  |                 +--+  +--------+                                  
  |  |                    |  |        |  +-- Potamilus_alatus **            
  |  |                    |  |        +--+                               
  |  |                    +--+           +-- Potamilus_streckersoni **      
  |  |                       |                                           
  |  |                       |  +----------- Leaunio_lienosus **            
  |  |                       |  |                                        
  |  |                       +--+  +-------- Venustaconcha_ellipsiformis ** 
  |  |                          |  |                                     
  |  |                          |  |     +-- Lampsilis_powellii **          
  |  |                          +--+-----+                               
  |  |                             |     +-- Lampsilis_siliquoidea **       
  |  |                             |                                     
  |  |                             |  +----- Lampsilis_ornata **            
  |  |                             +--+                                  
  |  |                                |  +-- Lampsilis_cardium **           
  |  |                                +--+                               
  |  |                                   +-- Lampsilis_cariosa **           
  |  |                                                                   
  +--+                                +----- Chamberlainia_somsakpanhai **  
     |           +--------------------+                                  
     |           |                    |  +-- Sinohyriopsis_cumingii **      
     |           |                    +--+                               
     |           |                       +-- Sinohyriopsis_schlegelii **    
     |           |                                                       
     |           |                       +-- Hyriopsis_bialata **           
     |           |                    +--+                               
     |  +--------+                    |  +-- Rectidens_sumatrensis **       
     |  |        |  +-----------------+                                  
     |  |        |  |                 |  +-- Lens_contradens **             
     |  |        |  |                 +--+                               
     |  |        |  |                    +-- Physunio_superbus **           
     |  |        |  |                                                    
     |  |        |  |           +----------- Potomida_littoralis **         
     |  |        +--+  +--------+                                        
     |  |           |  |        |  +-------- Pronodularia_japanensis **     
     |  |           |  |        +--+                                     
     |  |           |  |           |  +----- Lamprotula_leaii **            
     |  |           |  |           +--+                                  
     |  |           |  |              |  +-- Lamprotula_caveata **          
     |  |           +--+              +--+                               
     |  |              |                 +-- Lamprotula_cornuumlunae **     
     |  |              |                                                 
     |  |              |                 +-- Pseudodon_vondembuschianus **  
     |  |              |  +--------------+                               
     |  |              |  |              +-- Pilsbryoconcha_exilis **       
     |  |              +--+                                              
     |  |                 |  +-------------- Inversidens_rentianensis **    
     |  |                 |  |                                           
     +--+                 +--+           +-- Parvasolenaia_rivularis **     
        |                    |  +--------+                               
        |                    |  |        +-- Ptychorhynchus_pfisteri **     
        |                    +--+                                        
        |                       |  +-------- Microcondylaea_bonellii **     
        |                       +--+                                     
        |                          |  +----- Sinosolenaia_carinata **       
        |                          +--+                                  
        |                             |  +-- Sinosolenaia_iridinea **       
        |                             +--+                               
        |                                +-- Sinosolenaia_oleivora **       
        |                                                                
        |  +-------------------------------- Lamellidens_marginalis **      
        |  |                                                             
        |  |                             +-- Lepidodesma_aligera **         
        |  |  +--------------------------+                               
        |  |  |                          +-- Lepidodesma_languilati **      
        |  |  |                                                          
        |  |  |                    +-------- Aculamprotula_tortuosa **      
        |  |  |                    |                                     
        |  |  |     +--------------+     +-- Aculamprotula_coreana **       
        |  |  |     |              |  +--+                               
        +--+  |     |              |  |  +-- Aculamprotula_tientsinensis ** 
           |  |     |              +--+                                  
           |  |     |                 |  +-- Aculamprotula_polysticta **    
           |  |     |                 +--+                               
           |  |     |                    +-- Aculamprotula_scripta **       
           |  |     |                                                    
           |  |  +--+        +-------------- Unio_tumidus **                
           |  |  |  |  +-----+                                           
           |  |  |  |  |     |  +----------- Unio_crassus **                
           |  |  |  |  |     +--+                                        
           +--+  |  |  |        |  +-------- Unio_elongatulus **            
              |  |  |  |        +--+                                     
              |  |  |  |           |  +----- Unio_pictorum **               
              |  |  |  |           +--+                                  
              |  |  |  |              |  +-- Unio_delphinus **              
              |  |  +--+              +--+                               
              |  |     |                 +-- Unio_mancus **                 
              |  |     |                                                 
              |  |     |                 +-- Nodularia_breviconcha **       
              |  |     |  +--------------+                               
              |  |     |  |              +-- Nodularia_douglasiae **        
              |  |     |  |                                              
              |  |     |  |              +-- Pseudocuneopsis_capitata **    
              |  |     |  +--------------+                               
              |  |     +--+              +-- Pseudocuneopsis_sichuanensis **
              |  |        |                                              
              |  |        |              +-- Schistodesmus_lampreyanus **   
              +--+        |  +-----------+                               
                 |        |  |           +-- Schistodesmus_spinosus **      
                 |        +--+                                           
                 |           |  +----------- Tchangsinaia_piscicula **      
                 |           +--+                                        
                 |              |  +-------- Cuneopsis_rufescens **         
                 |              +--+                                     
                 |                 |  +----- Cuneopsis_celtiformis **       
                 |                 +--+                                  
                 |                    |  +-- Cuneopsis_demangei **          
                 |                    +--+                               
                 |                       +-- Cuneopsis_heudei **            
                 |                                                       
                 |                 +-------- Acuticosta_chinensis **        
                 |  +--------------+                                     
                 |  |              |  +----- Lanceolaria_gladiola **        
                 |  |              +--+                                  
                 |  |                 |  +-- Lanceolaria_grayii **          
                 |  |                 +--+                               
                 |  |                    +-- Lanceolaria_lanceolata **      
                 |  |                                                    
                 |  |        +-------------- Cristaria_plicata **           
                 +--+        |                                           
                    |  +-----+           +-- Anemina_arcaeformis **         
                    |  |     |  +--------+                               
                    |  |     |  |        +-- Anemina_euscaphys **           
                    |  |     +--+                                        
                    |  |        |  +-------- Beringiana_fukuharai **        
                    |  |        +--+                                     
                    |  |           |  +----- Sinanodonta_lucida **          
                    |  |           +--+                                  
                    +--+              |  +-- Sinanodonta_tumens **          
                       |              +--+                               
                       |                 +-- Sinanodonta_woodiana **        
                       |                                                 
                       |  +----------------- Anodonta_nuttalliana **        
                       |  |                                              
                       |  |        +-------- Pseudanodonta_complanata **    
                       |  |  +-----+                                     
                       +--+  |     |  +----- Anodonta_anatina **            
                          |  |     +--+                                  
                          |  |        |  +-- Anodonta_cygnea **             
                          |  |        +--+                               
                          +--+           +-- Anodonta_exulcerata **         
                             |                                           
                             |        +----- Pyganodon_grandis **           
                             |  +-----+                                  
                             |  |     |  +-- Utterbackia_imbecillis **      
                             |  |     +--+                               
                             +--+        +-- Utterbackia_peninsularis **    
                                |                                        
                                |  +-------- Lasmigona_complanata **        
                                +--+                                     
                                   |  +----- Prolasmidonta_heterodon **     
                                   +--+                                  
                                      |  +-- Alasmidonta_varicosa **        
                                      +--+                               
                                         +-- Platynaias_compressa **        

Bolotov et al., 2018 [Publ 3826], fig. 2

iT tT bi riS oT roT roS iD
85 85 50 4 0 0 0 106.0
                                   +-- Trapezidens_exolescens **     
                          +--------+                              
                          |        +-- Trapezidens_dolichorhynchus **
           +--------------+                                       
           |              |  +-------- Lamellidens_savadiensis **    
           |              +--+                                    
           |                 |  +----- Lamellidens_ferrugineus **    
           |                 +--+                                 
           |                    |  +-- Lamellidens_generosus **      
           |                    +--+                              
  +--------+                       +-- Lamellidens_mainwaringi **    
  |        |                                                      
  |        |                       +-- Parreysia_keralaensis **      
  |        |  +--------------------+                              
  |        |  |                    +-- Parreysia_nagpoorensis **     
  |        |  |                                                   
  |        |  |                 +----- Leoparreysia_olivacea **      
  |        +--+  +--------------+                                 
  |           |  |              |  +-- Leoparreysia_canefrii **      
  |           |  |              +--+                              
  |           |  |                 +-- Leoparreysia_tavoyensis **    
  |           |  |                                                
  |           +--+                 +-- Coelatura_aegyptiaca **       
  |              |  +--------------+                              
  |              |  |              +-- Nitia_teretiuscula **         
  |              |  |                                             
  |              |  |        +-------- Indonaia_shurtleffiana **     
  |              +--+  +-----+                                    
  |                 |  |     |  +----- Indonaia_rugosa **            
  |                 |  |     +--+                                 
  |                 |  |        |  +-- Indonaia_caerulea **          
  |                 |  |        +--+                              
  |                 +--+           +-- Indonaia_andersoniana **      
  |                    |                                          
  |                    |  +----------- Indochinella_pugio **         
  |                    |  |                                       
  |                    |  |     +----- Radiatula_myitkyinae **       
  |                    +--+  +--+                                 
  |                       |  |  |  +-- Radiatula_mouhoti **          
  |                       |  |  +--+                              
  |                       +--+     +-- Radiatula_chaudhurii **       
  |                          |                                    
  |                          |  +----- Scabies_mandarinus **         
  |                          +--+                                 
  |                             |  +-- Scabies_humilis **            
  |                             +--+                              
--+                                +-- Scabies_phaselus **           
  |                                                               
  |                                +-- Indopseudodon_indawgyiensis **
  |                                |                              
  |                             +--+-- Indopseudodon_bogani **       
  |                             |  |                              
  |                             |  +-- Indopseudodon_salwenianus **  
  |                             |                                 
  |                             |  +-- Bineurus_mouhotii **          
  |  +--------------------------+  |                              
  |  |                          |  +-- Pseudodon_vagulus **          
  |  |                          |  |                              
  |  |                          |  +-- Pseudodon_mekongi **          
  |  |                          |  |                              
  |  |                          |  +-- Pilsbryoconcha_exilis **      
  |  |                          +--+                              
  |  |                             +-- Pilsbryoconcha_carinifera **  
  |  |                             |                              
  |  |                             +-- Bineurus_loeiensis **         
  |  |                             |                              
  |  |                             +-- Bineurus_exilis **            
  |  |                             |                              
  |  |                             +-- Pseudodon_vondembuschianus ** 
  |  |                                                            
  |  |                             +-- Ensidens_telus **             
  |  |                             |                              
  |  |                             +-- Ensidens_ingallsianus **      
  |  |                             |                              
  |  |                             +-- Ensidens_jaculus **           
  |  |                             |                              
  |  |                             +-- Hyriopsis_desowitzi **        
  |  |                          +--+                              
  |  |                          |  +-- Hyriopsis_myersiana **        
  |  |                          |  |                              
  |  |                          |  +-- Hyriopsis_khoratensis **      
  |  |                          |  |                              
  +--+                          |  +-- Ensidens_spiculus **          
     |                          |  |                              
     |                          |  +-- Ensidens_sagittarius **       
     |                          |                                 
     |  +-----------------------+  +-- Lens_eximius **               
     |  |                       |  |                              
     |  |                       |  +-- Physunio_modelli[1] **        
     |  |                       |  |                              
     |  |                       |  +-- Yaukthwa_nesemanni **         
     |  |                       |  |                              
     |  |                       |  +-- Yaukthwa_panhai **            
     |  |                       |  |                              
     |  |                       |  +-- Lens_comptus **               
     |  |                       |  |                              
     |  |                       +--+-- Lens_novoselovi **            
     |  |                          |                              
     |  |                          +-- Physunio_modelli[2] **        
     |  |                          |                              
     |  |                          +-- Physunio **                   
     |  |                          |                              
     |  |                          +-- Yaukthwa_inlenensis **        
     |  |                          |                              
     |  |                          +-- Lens_contradens **            
     |  |                          |                              
     |  |                          +-- Lens_micropterus **           
     |  |                                                         
     |  |                          +-- Lamprotula_leaii **           
     +--+                          |                              
        |                          +-- Potomida_littoralis **        
        |                          |                              
        |                          +-- Pronodularia_japanensis **    
        |                          |                              
        |                          +-- Gonidea_angulata **           
        |                          |                              
        |                          +-- Leguminaia_wheatleyi **       
        |                          |                              
        |                          +-- Microcondylaea_bonellii **    
        |                          |                              
        |  +-----------------------+-- Amblema_plicata **            
        |  |                       |                              
        |  |                       +-- Ortmanniana_ligamentina **    
        |  |                       |                              
        |  |                       +-- Lampsilis_cardium **          
        |  |                       |                              
        |  |                       +-- Cambarunio_iris **            
        |  |                       |                              
        |  |                       +-- Elliptio_complanata **        
        |  |                       |                              
        |  |                       +-- Pleurobema_sintoxia **        
        |  |                       |                              
        +--+                       +-- Quadrula_quadrula **          
           |                                                      
           |                    +----- Unio_tumidus **               
           |                 +--+                                 
           |                 |  |  +-- Unio_crassus **               
           |                 |  +--+                              
           |  +--------------+     +-- Unio_pictorum **              
           |  |              |                                    
           |  |              |  +----- Nodularia_douglasiae[1] **    
           |  |              +--+                                 
           |  |                 |  +-- Nodularia_nuxpersicae **      
           |  |                 +--+                              
           +--+                    +-- Nodularia_douglasiae[2] **    
              |                                                   
              |                    +-- Lanceolaria_gladiola **       
              |  +-----------------+                              
              |  |                 +-- Lanceolaria_grayii **         
              |  |                                                
              |  |           +-------- Cristaria_plicata **          
              +--+  +--------+                                    
                 |  |        |  +----- Pletholophus_tenuis **        
                 |  |        +--+                                 
                 |  |           |  +-- Sinanodonta_jourdyi **        
                 +--+           +--+                              
                    |              +-- Sinanodonta_lauta **          
                    |                                             
                    |  +-------------- Anodonta_anatina **           
                    +--+                                          
                       |  +----------- Pyganodon_grandis **          
                       +--+                                       
                          |  +-------- Simpsonaias_ambigua **        
                          +--+                                    
                             |  +----- Strophitus_undulatus **       
                             +--+                                 
                                |  +-- Alasmidonta_marginata **      
                                +--+                              
                                   +-- Lasmigona_costata **          

Wu et al., 2023 [Publ 4339], fig. 4, S5-aa

iT tT bi riS oT roT roS iD
85 85 83 4 11 7 1 121.0
                                         +-- Gibbosula_crassa            
                             +-----------+                               
                             |           +-- Gibbosula_rochechouartii    
  +--------------------------+                                           
  |                          |  +----------- Cumberlandia_monodonta      
  |                          +--+                                        
  |                             |  +-------- Pseudunio_marocanus         
  |                             +--+                                     
  |                                |  +----- Margaritifera_falcata       
  |                                +--+                                  
  |                                   |  +-- Margaritifera_dahurica      
  |                                   +--+                               
  |                                      +-- Margaritifera_margaritifera 
  |                                                                      
  |                                      +-- Quadrula_quadrula **           
  |           +--------------------------+                               
  |           |                          +-- Uniomerus_tetralasmus **       
  |           |                                                          
  |  +--------+                          +-- Elliptio_complanata **         
  |  |        |  +-----------------------+                               
  |  |        |  |                       +-- Pleurobema_oviforme **         
  |  |        +--+                                                       
  |  |           |  +----------------------- Amblema_plicata **             
  |  |           |  |                                                    
  |  |           +--+  +-------------------- Popenaias_popeii **            
--+  |              |  |                                                 
  |  |              +--+  +----------------- Toxolasma_parvum **            
  |  |                 |  |                                              
  |  |                 |  |           +----- Potamilus_leptodon **          
  |  |                 +--+  +--------+                                  
  |  |                    |  |        |  +-- Potamilus_alatus **            
  |  |                    |  |        +--+                               
  |  |                    |  |           +-- Potamilus_streckersoni **      
  |  |                    +--+                                           
  |  |                       |  +----------- Leaunio_lienosus **            
  |  |                       |  |                                        
  |  |                       |  |     +----- Lampsilis_ornata **            
  |  |                       +--+  +--+                                  
  |  |                          |  |  |  +-- Lampsilis_cardium **           
  |  |                          |  |  +--+                               
  |  |                          +--+     +-- Lampsilis_cariosa **           
  |  |                             |                                     
  |  |                             |  +----- Venustaconcha_ellipsiformis ** 
  |  |                             +--+                                  
  |  |                                |  +-- Lampsilis_powellii **          
  |  |                                +--+                               
  |  |                                   +-- Lampsilis_siliquoidea **       
  +--+                                                                   
     |                                +----- Chamberlainia_somsakpanhai **  
     |              +-----------------+                                  
     |              |                 |  +-- Sinohyriopsis_cumingii **      
     |              |                 +--+                               
     |              |                    +-- Sinohyriopsis_schlegelii **    
     |              |                                                    
     |              |                    +-- Hyriopsis_bialata **           
     |              |                 +--+                               
     |  +-----------+                 |  +-- Rectidens_sumatrensis **       
     |  |           |  +--------------+                                  
     |  |           |  |              |  +-- Lens_contradens **             
     |  |           |  |              +--+                               
     |  |           |  |                 +-- Physunio_superbus **           
     |  |           |  |                                                 
     |  |           |  |        +----------- Potomida_littoralis **         
     |  |           +--+  +-----+                                        
     |  |              |  |     |  +-------- Pronodularia_japanensis **     
     |  |              |  |     +--+                                     
     |  |              |  |        |  +----- Lamprotula_leaii **            
     |  |              |  |        +--+                                  
     |  |              |  |           |  +-- Lamprotula_caveata **          
     |  |              +--+           +--+                               
     |  |                 |              +-- Lamprotula_cornuumlunae **     
     |  |                 |                                              
     |  |                 |              +-- Pseudodon_vondembuschianus **  
     |  |                 |  +-----------+                               
     |  |                 |  |           +-- Pilsbryoconcha_exilis **       
     |  |                 +--+                                           
     |  |                    |  +----------- Microcondylaea_bonellii **     
     +--+                    |  |                                        
        |                    +--+        +-- Parvasolenaia_rivularis **     
        |                       |  +-----+                               
        |                       |  |     +-- Ptychorhynchus_pfisteri **     
        |                       +--+                                     
        |                          |  +----- Sinosolenaia_carinata **       
        |                          +--+                                  
        |                             |  +-- Sinosolenaia_iridinea **       
        |                             +--+                               
        |                                +-- Sinosolenaia_oleivora **       
        |                                                                
        |  +-------------------------------- Lamellidens_marginalis **      
        |  |                                                             
        |  |                             +-- Lepidodesma_aligera **         
        |  |  +--------------------------+                               
        |  |  |                          +-- Lepidodesma_languilati **      
        |  |  |                                                          
        |  |  |                    +-------- Aculamprotula_tortuosa **      
        |  |  |                    |                                     
        |  |  |     +--------------+     +-- Aculamprotula_coreana **       
        |  |  |     |              |  +--+                               
        +--+  |     |              |  |  +-- Aculamprotula_tientsinensis ** 
           |  |     |              +--+                                  
           |  |     |                 |  +-- Aculamprotula_polysticta **    
           |  |     |                 +--+                               
           |  |     |                    +-- Aculamprotula_scripta **       
           |  |     |                                                    
           |  |  +--+        +-------------- Unio_tumidus **                
           |  |  |  |  +-----+                                           
           |  |  |  |  |     |  +----------- Unio_crassus **                
           |  |  |  |  |     +--+                                        
           +--+  |  |  |        |  +-------- Unio_elongatulus **            
              |  |  |  |        +--+                                     
              |  |  |  |           |  +----- Unio_pictorum **               
              |  |  |  |           +--+                                  
              |  |  |  |              |  +-- Unio_delphinus **              
              |  |  +--+              +--+                               
              |  |     |                 +-- Unio_mancus **                 
              |  |     |                                                 
              |  |     |                 +-- Nodularia_breviconcha **       
              |  |     |              +--+                               
              |  |     |              |  +-- Nodularia_nipponensis **       
              |  |     |  +-----------+                                  
              |  |     |  |           |  +-- Pseudocuneopsis_capitata **    
              |  |     |  |           +--+                               
              |  |     +--+              +-- Pseudocuneopsis_sichuanensis **
              |  |        |                                              
              |  |        |  +-------------- Tchangsinaia_piscicula **      
              |  |        |  |                                           
              +--+        +--+           +-- Schistodesmus_lampreyanus **   
                 |           |  +--------+                               
                 |           |  |        +-- Schistodesmus_spinosus **      
                 |           +--+                                        
                 |              |  +-------- Cuneopsis_rufescens **         
                 |              +--+                                     
                 |                 |  +----- Cuneopsis_celtiformis **       
                 |                 +--+                                  
                 |                    |  +-- Cuneopsis_demangei **          
                 |                    +--+                               
                 |                       +-- Cuneopsis_heudei **            
                 |                                                       
                 |                 +-------- Acuticosta_chinensis **        
                 |        +--------+                                     
                 |        |        |  +----- Lanceolaria_gladiola **        
                 |        |        +--+                                  
                 |        |           |  +-- Lanceolaria_grayii **          
                 |        |           +--+                               
                 |        |              +-- Lanceolaria_lanceolata **      
                 |        |                                              
                 |        |           +----- Cristaria_plicata **           
                 |        |     +-----+                                  
                 |        |     |     |  +-- Anemina_arcaeformis **         
                 +--------+     |     +--+                               
                          |  +--+        +-- Anemina_euscaphys **           
                          |  |  |                                        
                          |  |  |  +-------- Beringiana_fukuharai **        
                          |  |  +--+                                     
                          |  |     |  +----- Sinanodonta_lucida **          
                          |  |     +--+                                  
                          |  |        |  +-- Sinanodonta_tumens **          
                          |  |        +--+                               
                          |  |           +-- Sinanodonta_woodiana **        
                          +--+                                           
                             |           +-- Anodonta_nuttalliana **        
                             |     +-----+                               
                             |     |     +-- Pseudanodonta_complanata **    
                             |  +--+                                     
                             |  |  |  +----- Anodonta_anatina **            
                             |  |  +--+                                  
                             |  |     |  +-- Anodonta_cygnea **             
                             |  |     +--+                               
                             |  |        +-- Anodonta_exulcerata **         
                             +--+                                        
                                |     +----- Pyganodon_grandis **           
                                |  +--+                                  
                                |  |  |  +-- Utterbackia_peninsularis **    
                                |  |  +--+                               
                                |  |     +-- Utterbackia_imbecillis **      
                                +--+                                     
                                   |     +-- Alasmidonta_varicosa **        
                                   |  +--+                               
                                   |  |  +-- Platynaias_compressa **        
                                   +--+                                  
                                      |  +-- Prolasmidonta_heterodon **     
                                      +--+                               
                                         +-- Lasmigona_complanata **        

Konopleva et al., 2023 [Publ 4262], fig. 8

iT tT bi riS oT roT roS iD
84 84 80 2 0 0 0 120.0
                 +----------------------- Indonaia_andersoniana **        
                 |                                                     
  +--------------+  +-------------------- Indochinella_pugio **           
  |              |  |                                                  
  |              +--+  +----------------- Radiatula_mouhoti **            
  |                 |  |                                               
  |                 +--+  +-------------- Scabiellus_songkramensis **     
  |                    |  |                                            
  |                    |  |           +-- Unionetta_fabagina **           
  |                    +--+  +--------+                                
  |                       |  |        +-- Harmandia_somboriensis **       
  |                       |  |                                         
  |                       |  |        +-- Scabies_humilis **              
  |                       +--+  +-----+                                
  |                          |  |     +-- Scabies_pilata **               
  |                          |  |                                      
  |                          |  |  +----- Scabies_crispata **             
  |                          |  +--+                                   
  |                          +--+  |  +-- Scabies_nucleus **              
  |                             |  +--+                                
  |                             |     +-- Scabies_mandarinus **           
  |                             |                                      
  |                             |     +-- Scabies_phaselus **             
  |                             |     |                                
  |                             +-----+-- Scabies_anceps **               
  |                                   |                                
  |                                   +-- Scabies_scobinatus **           
  |                                                                    
  |                             +-------- Indopseudodon_kayinensis **     
  |     +-----------------------+                                      
  |     |                       |  +----- Indopseudodon_indawgyiensis **  
  |     |                       +--+                                   
  |     |                          |  +-- Indopseudodon_salwenianus **    
  |     |                          +--+                                
  |     |                             +-- Indopseudodon_bogani **         
  |     |                                                              
  |     |                             +-- Pilsbryoconcha_schomburgki **   
--+  +--+                    +--------+                                
  |  |  |                    |        +-- Pilsbryoconcha_exilis[1] **     
  |  |  |  +-----------------+                                         
  |  |  |  |                 |  +-------- Pilsbryoconcha **               
  |  |  |  |                 +--+                                      
  |  |  |  |                    |  +----- Pilsbryoconcha_linguaeformis ** 
  |  |  |  |                    +--+                                   
  |  |  |  |                       |  +-- Pilsbryoconcha_exilis[2] **     
  |  |  +--+                       +--+                                
  |  |     |                          +-- Pilsbryoconcha_carinifera **    
  |  |     |                                                           
  |  |     |  +-------------------------- Songkhlanaia_tamodienica **     
  |  |     |  |                                                        
  |  |     |  |                       +-- Namkongnaia_inkhavilayi **      
  |  |     |  |  +--------------------+                                
  |  |     +--+  |                    +-- Namkongnaia_lemeslei **         
  |  |        |  |                                                     
  |  |        |  |              +-------- Pseudodon_lenyanensis **        
  |  |        |  |  +-----------+                                      
  |  |        |  |  |           |  +----- Pseudodon_vagulus **            
  |  |        +--+  |           +--+                                   
  |  |           |  |              |  +-- Pseudodon_vondembuschianus **   
  |  |           |  |              +--+                                
  |  |           |  |                 +-- Pseudodon_mekongi **            
  |  |           |  |                                                  
  |  |           |  |                 +-- Sundadontina_taskaevi **        
  |  |           +--+        +--------+                                
  |  |              |        |        +-- Sundadontina_brandti **         
  |  |              |  +-----+                                         
  |  |              |  |     |  +-------- Sundadontina_tumida **          
  |  |              |  |     +--+                                      
  |  |              |  |        |  +----- Sundadontina_tanintharyiensis **
  |  |              |  |        +--+                                   
  |  |              |  |           |  +-- Sundadontina_plugpomenica **    
  |  |              +--+           +--+                                
  |  |                 |              +-- Sundadontina_cumingii **        
  +--+                 |                                               
     |                 |  +-------------- Nyeinchanconcha_nyeinchani **   
     |                 |  |                                            
     |                 |  |        +----- Thaiconcha_thaiensis **         
     |                 +--+  +-----+                                   
     |                    |  |     |  +-- Thaiconcha_munelliptica **      
     |                    |  |     +--+                                
     |                    +--+        +-- Thaiconcha_callifera **         
     |                       |                                         
     |                       |  +-------- Bineurus_loeiensis **           
     |                       +--+                                      
     |                          |  +----- Bineurus_mouhotii **            
     |                          +--+                                   
     |                             |  +-- Bineurus_anodontinum **         
     |                             +--+                                
     |                                +-- Bineurus_exilis **              
     |                                                                 
     |                                +-- Gonidea_angulata **             
     |                             +--+                                
     |                             |  +-- Leguminaia_wheatleyi **         
     |     +-----------------------+                                   
     |     |                       |  +-- Lamprotula_leaii **             
     |     |                       +--+                                
     |     |                          +-- Potomida_littoralis **          
     |     |                                                           
     |     |                          +-- Ensidens_ingallsianus **        
     |     |                    +-----+                                
     |     |                    |     +-- Ensidens_telus **               
     |     |                    |                                      
     |     |     +--------------+     +-- Ensidens_jaculus **             
     |     |     |              |  +--+                                
     |     |     |              |  |  +-- Ensidens_spiculus[1] **         
     |     |     |              +--+                                   
     |     |     |                 |  +-- Ensidens_spiculus[2] **         
     |     |  +--+                 +--+                                
     |     |  |  |                    +-- Ensidens_dugasti **             
     +-----+  |  |                                                     
           |  |  |                    +-- Hyriopsis_desowitzi **          
           |  |  |  +-----------------+                                
           |  |  |  |                 +-- Hyriopsis_delaportei **         
           |  |  +--+                                                  
           |  |     |  +----------------- Rectidens_sumatrensis **        
           |  |     |  |                                               
           |  |     +--+  +-------------- Hyriopsis_panhai **             
           |  |        |  |                                            
           |  |        +--+  +----------- Hyriopsis_myersiana **          
           |  |           |  |                                         
           |  |           +--+        +-- Hyriopsis_khoratensis **        
           |  |              |  +-----+                                
           |  |              |  |     +-- Hyriopsis_bialata **            
           |  |              +--+                                      
           |  |                 |  +----- Hyriopsis_sakhonensis **        
           +--+                 +--+                                   
              |                    |  +-- Hyriopsis_phuphaniensis **      
              |                    +--+                                
              |                       +-- Hyriopsis_kratiensis **         
              |                                                        
              |                    +----- Physunio_modelli **             
              |              +-----+                                   
              |              |     |  +-- Physunio_superbus **            
              |              |     +--+                                
              |              |        +-- Physunio_massini **             
              |        +-----+                                         
              |        |     |        +-- Trapezoideus_thachiadensis **   
              |        |     |  +-----+                                
              |        |     |  |     +-- Trapezoideus_subviridis **      
              |        |     +--+                                      
              |        |        |  +----- Trapezoideus_mitanensis **      
              |        |        +--+                                   
              |        |           |  +-- Trapezoideus_foliaceus **       
              +--------+           +--+                                
                       |              +-- Trapezoideus_lenya **           
                       |                                               
                       |              +-- Yaukthwa_nesemanni **           
                       |  +-----------+                                
                       |  |           +-- Yaukthwa_elongatula **          
                       |  |                                            
                       |  |        +----- Lens_misellus **                
                       +--+  +-----+                                   
                          |  |     |  +-- Lens_contradens **              
                          |  |     +--+                                
                          |  |        +-- Lens_micropterus **             
                          +--+                                         
                             |     +----- Lens_pallegoixi **              
                             |  +--+                                   
                             |  |  |  +-- Lens_novoselovi **              
                             |  |  +--+                                
                             +--+     +-- Lens_comptus **                 
                                |                                      
                                |  +----- Lens_maenamensis **             
                                +--+                                   
                                   |  +-- Lens_eximius **                 
                                   +--+                                
                                      +-- Lens_rolfbrandti **             

Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2

iT tT bi riS oT roT roS iD
74 74 64 1 0 0 0 111.0
                                         +-- Uniomerus_declivis **         
                       +-----------------+                              
                       |                 +-- Uniomerus_tetralasmus **      
  +--------------------+                                                
  |                    |  +----------------- Theliderma_metanevra **       
  |                    |  |                                             
  |                    +--+  +-------------- Quadrula_verrucosa **         
  |                       |  |                                          
  |                       |  |           +-- Quadrula_quadrula **          
  |                       +--+  +--------+                              
  |                          |  |        +-- Quadrula_nobilis **           
  |                          |  |                                       
  |                          +--+        +-- Pustulosa_infucata **         
  |                             |  +-----+                              
  |                             |  |     +-- Pustulosa_kleiniana **        
  |                             +--+                                    
  |                                |  +----- Cyclonaias_tuberculata **     
  |                                +--+                                 
  |                                   |  +-- Pustulosa_pustulosa **        
  |                                   +--+                              
  |                                      +-- Pustulosa_succissa **         
  |                                                                     
  |                                   +----- Plectomerus_dombeyanus **     
  |                 +-----------------+                                 
  |                 |                 |  +-- Reginaia_ebenus **            
  |                 |                 +--+                              
--+                 |                    +-- Reginaia_rotulata **          
  |                 |                                                   
  |                 |                 +----- Popenaias_popeii **           
  |  +--------------+  +--------------+                                 
  |  |              |  |              |  +-- Amblema_elliottii **          
  |  |              |  |              +--+                              
  |  |              |  |                 +-- Amblema_plicata **            
  |  |              |  |                                                
  |  |              |  |  +----------------- Potamilus_alatus **           
  |  |              +--+  |                                             
  |  |                 |  +----------------- Potamilus_capax **            
  |  |                 |  |                                             
  |  |                 |  +----------------- Glebula_rotundata **          
  |  |                 |  |                                             
  |  |                 |  |              +-- Toxolasma_lividum **          
  |  |                 |  +--------------+                              
  |  |                 +--+              +-- Paetulunio_fabalis **         
  |  |                    |                                             
  |  |                    |              +-- Dromus_dromas **              
  |  |                    +--------------+                              
  |  |                    |              +-- Lemiox_rimosus **             
  |  |                    |                                             
  |  |                    |              +-- Lampsilis_ovata **            
  +--+                    |  +-----------+                              
     |                    |  |           +-- Obliquaria_reflexa **         
     |                    +--+                                          
     |                       |  +----------- Lampsilis_virescens **        
     |                       |  |                                       
     |                       +--+        +-- Epioblasma_rangiana **        
     |                          |  +-----+                              
     |                          |  |     +-- Obovaria_arkansasensis **     
     |                          +--+                                    
     |                             |  +----- Obovaria_olivaria **          
     |                             +--+                                 
     |                                |  +-- Venustaconcha_ellipsiformis **
     |                                +--+                              
     |                                   +-- Venustaconcha_pleasii **      
     |                                                                  
     |  +----------------------------------- Elliptoideus_sloatianus **    
     |  |                                                               
     |  |  +-------------------------------- Pleurobema_stabile **         
     |  |  |                                                            
     |  |  |                             +-- Pleurobema_clava **           
     |  |  |                 +-----------+                              
     +--+  |                 |           +-- Pleurobema_oviforme **        
        |  |                 |                                          
        |  |  +--------------+  +----------- Pleurobema_pyriforme **       
        |  |  |              |  |                                       
        |  |  |              |  |        +-- Pleurobema_athearni **        
        |  |  |              |  |     +--+                              
        +--+  |              +--+     |  +-- Pleurobema_georgianum **      
           |  |                 |  +--+                                 
           |  |                 |  |  |  +-- Pleurobema_decisum **         
           |  |                 |  |  +--+                              
           |  |                 |  |     +-- Pleurobema_hanleyianum **     
           |  |                 +--+                                    
           |  |                    |  +----- Pleurobema_beadleianum **     
           |  |                    |  |                                 
           |  |                    |  +----- Pleurobema_strodeanum **      
           |  |                    +--+                                 
           |  |                       +----- Pleurobema_rubellum **        
           +--+                       |                                 
              |                       |  +-- Pleurobema_perovatum **       
              |                       +--+                              
              |                          +-- Pleurobema_taitianum **       
              |                                                         
              |                    +-------- Pleurobema_riddellii **       
              |  +-----------------+                                    
              |  |                 |  +----- Pleurobema_cordatum **        
              |  |                 +--+                                 
              |  |                    |  +-- Pleurobema_rubrum **          
              |  |                    +--+                              
              |  |                       +-- Pleurobema_sintoxia **        
              |  |                                                      
              |  |        +----------------- Fusconaia_subrotunda **       
              |  |        |                                             
              +--+  +-----+              +-- Fusconaia_cor **              
                 |  |     |  +-----------+                              
                 |  |     |  |           +-- Fusconaia_cuneolus **         
                 |  |     +--+                                          
                 |  |        |  +----------- Fusconaia_ozarkensis **       
                 |  |        |  |                                       
                 |  |        +--+        +-- Fusconaia_escambia **         
                 |  |           |  +-----+                              
                 |  |           |  |     +-- Fusconaia_burkei **           
                 |  |           +--+                                    
                 +--+              |  +----- Fusconaia_masoni **           
                    |              +--+                                 
                    |                 |  +-- Fusconaia_flava **            
                    |                 +--+                              
                    |                    +-- Fusconaia_cerina **           
                    |                                                   
                    |                 +----- Pleuronaia_gibber **          
                    |  +--------------+                                 
                    |  |              |  +-- Pleuronaia_barnesiana **      
                    |  |              +--+                              
                    |  |                 +-- Pleuronaia_dolabelloides **   
                    +--+                                                
                       |                 +-- Eurynia_dilatata **           
                       |  +--------------+                              
                       |  |              +-- Parvaspina_collina **         
                       |  |                                             
                       +--+              +-- Hemistena_lata **             
                          |  +-----------+                              
                          |  |           +-- Plethobasus_cyphyus **        
                          |  |                                          
                          +--+  +----------- Elliptio_shepardiana **       
                             |  |                                       
                             |  |  +-------- Elliptio_complanata **        
                             +--+  |                                    
                                |  |  +----- Elliptio_purpurella **        
                                |  +--+                                 
                                |  |  |  +-- Elliptio_arca **              
                                +--+  +--+                              
                                   |     +-- Elliptio_arctata **           
                                   |                                    
                                   |     +-- Elliptio_mcmichaeli **        
                                   |     |                              
                                   +-----+-- Elliptio_pullata **           
                                         |                              
                                         +-- Elliptio_crassidens **        

Campbell & Lydeard, 2012 [Publ 3309], fig. 3

iT tT bi riS oT roT roS iD
74 74 68 1 0 0 0 109.0
                                      +-- Uniomerus_declivis **         
                       +--------------+                              
                       |              +-- Uniomerus_tetralasmus **      
                       |                                             
  +--------------------+              +-- Quadrula_verrucosa **         
  |                    |  +-----------+                              
  |                    |  |           +-- Quadrula_nobilis **           
  |                    |  |                                          
  |                    +--+  +----------- Quadrula_quadrula **          
  |                       |  |                                       
  |                       |  +----------- Theliderma_metanevra **       
  |                       +--+                                       
  |                          |        +-- Pustulosa_infucata **         
  |                          |  +-----+                              
  |                          |  |     +-- Pustulosa_kleiniana **        
  |                          +--+                                    
  |                             |  +----- Cyclonaias_tuberculata **     
  |                             +--+                                 
  |                                |  +-- Pustulosa_pustulosa **        
  |                                +--+                              
  |                                   +-- Pustulosa_succissa **         
  |                                                                  
  |                                   +-- Amblema_elliottii **          
  |                                +--+                              
  |                                |  +-- Amblema_plicata **            
  |     +--------------------------+                                 
  |     |                          |  +-- Reginaia_ebenus **            
--+     |                          +--+                              
  |     |                             +-- Reginaia_rotulata **          
  |  +--+                                                            
  |  |  |  +----------------------------- Popenaias_popeii **           
  |  |  |  |                                                         
  |  |  |  |                          +-- Glebula_rotundata **          
  |  |  +--+  +-----------------------+                              
  |  |     |  |                       +-- Paetulunio_fabalis **         
  |  |     |  |                                                      
  |  |     +--+                       +-- Toxolasma_lividum **          
  |  |        |  +--------------------+                              
  |  |        |  |                    +-- Plectomerus_dombeyanus **     
  |  |        |  |                                                   
  |  |        +--+  +-------------------- Obliquaria_reflexa **         
  |  |           |  |                                                
  |  |           |  |                 +-- Dromus_dromas **              
  |  |           |  |              +--+                              
  |  |           +--+              |  +-- Lemiox_rimosus **             
  |  |              |  +-----------+                                 
  |  |              |  |           |  +-- Potamilus_alatus **           
  |  |              |  |           +--+                              
  |  |              +--+              +-- Potamilus_capax **            
  |  |                 |                                             
  |  |                 |  +-------------- Lampsilis_ovata **            
  +--+                 |  |                                          
     |                 +--+  +----------- Lampsilis_virescens **        
     |                    |  |                                       
     |                    +--+        +-- Epioblasma_rangiana **        
     |                       |  +-----+                              
     |                       |  |     +-- Obovaria_arkansasensis **     
     |                       +--+                                    
     |                          |  +----- Obovaria_olivaria **          
     |                          +--+                                 
     |                             |  +-- Venustaconcha_ellipsiformis **
     |                             +--+                              
     |                                +-- Venustaconcha_pleasii **      
     |                                                               
     |  +-------------------------------- Elliptoideus_sloatianus **    
     |  |                                                            
     |  |                             +-- Eurynia_dilatata **           
     |  |  +--------------------------+                              
     |  |  |                          +-- Parvaspina_collina **         
     |  |  |                                                         
     |  |  |                          +-- Hemistena_lata **             
     |  |  +--------------------------+                              
     |  |  |                          +-- Plethobasus_cyphyus **        
     +--+  |                                                         
        |  |                 +----------- Elliptio_shepardiana **       
        |  |                 |                                       
        |  |                 |     +----- Elliptio_crassidens **        
        |  +-----------------+  +--+                                 
        |  |                 |  |  |  +-- Elliptio_mcmichaeli **        
        |  |                 |  |  +--+                              
        |  |                 +--+     +-- Elliptio_pullata **           
        |  |                    |                                    
        |  |                    |  +----- Elliptio_complanata **        
        +--+                    |  |                                 
           |                    +--+  +-- Elliptio_purpurella **        
           |                       |  |                              
           |                       +--+-- Elliptio_arca **              
           |                          |                              
           |                          +-- Elliptio_arctata **           
           |                                                         
           |                       +----- Pleuronaia_gibber **          
           |  +--------------------+                                 
           |  |                    |  +-- Pleuronaia_barnesiana **      
           |  |                    +--+                              
           |  |                       +-- Pleuronaia_dolabelloides **   
           |  |                                                      
           |  |           +-------------- Fusconaia_ozarkensis **       
           |  |           |                                          
           |  |           |        +----- Fusconaia_subrotunda **       
           |  |  +--------+  +-----+                                 
           +--+  |        |  |     |  +-- Fusconaia_cor **              
              |  |        |  |     +--+                              
              |  |        |  |        +-- Fusconaia_cuneolus **         
              |  |        +--+                                       
              |  |           |        +-- Fusconaia_escambia **         
              |  |           |  +-----+                              
              |  |           |  |     +-- Fusconaia_burkei **           
              |  |           +--+                                    
              |  |              |  +----- Fusconaia_masoni **           
              +--+              +--+                                 
                 |                 |  +-- Fusconaia_cerina **           
                 |                 +--+                              
                 |                    +-- Fusconaia_flava **            
                 |                                                   
                 |              +-------- Pleurobema_riddellii **       
                 |  +-----------+                                    
                 |  |           |  +----- Pleurobema_cordatum **        
                 |  |           +--+                                 
                 |  |              |  +-- Pleurobema_rubrum **          
                 |  |              +--+                              
                 +--+                 +-- Pleurobema_sintoxia **        
                    |                                                
                    |  +----------------- Pleurobema_stabile **         
                    |  |                                             
                    |  |     +----------- Pleurobema_strodeanum **      
                    |  |  +--+                                       
                    +--+  |  |  +-------- Pleurobema_rubellum **        
                       |  |  +--+                                    
                       |  |     |  +----- Pleurobema_beadleianum **     
                       |  |     +--+                                 
                       |  |        |  +-- Pleurobema_perovatum **       
                       +--+        +--+                              
                          |           +-- Pleurobema_taitianum **       
                          |                                          
                          |           +-- Pleurobema_clava **           
                          |  +--------+                              
                          |  |        +-- Pleurobema_oviforme **        
                          +--+                                       
                             |  +-------- Pleurobema_pyriforme **       
                             |  |                                    
                             +--+     +-- Pleurobema_athearni **        
                                |  +--+                              
                                |  |  +-- Pleurobema_georgianum **      
                                +--+                                 
                                   |  +-- Pleurobema_decisum **         
                                   +--+                              
                                      +-- Pleurobema_hanleyianum **     

Huang et al., 2019 [Publ 3843], fig. S1

iT tT bi riS oT roT roS iD
73 73 55 2 2 2 1 100.0
                                      +-- Gibbosula_rochechouartii   
  +-----------------------------------+                              
  |                                   +-- Margaritifera_dahurica     
  |                                                                  
  |                                +----- Sinohyriopsis_cumingii[2] **  
  |                 +--------------+                                 
  |                 |              |  +-- Sinohyriopsis_cumingii[1] **  
  |                 |              +--+                              
  |                 |                 +-- Sinohyriopsis_schlegelii **   
  |                 |                                                
  |                 |              +----- Parvasolenaia_rivularis **    
  |  +--------------+  +-----------+                                 
  |  |              |  |           |  +-- Parvasolenaia_triangularis ** 
  |  |              |  |           +--+                              
  |  |              |  |              +-- Ptychorhynchus_pfisteri **    
  |  |              |  |                                             
  |  |              |  |        +-------- Sinosolenaia_recognita **     
  |  |              +--+        |                                    
--+  |                 |  +-----+  +----- Sinosolenaia_oleivora[2] **   
  |  |                 |  |     |  |                                 
  |  |                 |  |     +--+  +-- Sinosolenaia_oleivora[1] **   
  |  |                 |  |        |  |                              
  |  |                 |  |        +--+-- Sinosolenaia_iridinea **      
  |  |                 +--+           |                              
  |  |                    |           +-- Sinosolenaia_oleivora[3] **   
  |  |                    |                                          
  |  |                    |  +----------- Sinosolenaia_carinata **      
  |  |                    |  |                                       
  |  |                    +--+  +-------- Lamprotula_leaii **           
  |  |                       |  |                                    
  |  |                       +--+  +----- Lamprotula_cornuumlunae[1] ** 
  |  |                          |  |                                 
  |  |                          |  +----- Lamprotula_cornuumlunae[3] ** 
  |  |                          +--+                                 
  |  |                             +----- Lamprotula_cornuumlunae[4] ** 
  +--+                             |                                 
     |                             |  +-- Lamprotula_cornuumlunae[2] ** 
     |                             +--+                              
     |                                +-- Lamprotula_caveata **         
     |                                                               
     |                    +-------------- Anemina_arcaeformis[2] **     
     |                    |                                          
     |  +-----------------+  +----------- Anemina_arcaeformis[5] **     
     |  |                 |  |                                       
     |  |                 |  |  +-------- Anemina_arcaeformis[3] **     
     |  |                 +--+  |                                    
     |  |                    |  +-------- Anemina_arcaeformis[7] **     
     |  |                    |  |                                    
     |  |                    +--+  +----- Anemina_arcaeformis[4] **     
     |  |                       |  |                                 
     |  |                       |  +----- Anemina_arcaeformis[6] **     
     |  |                       |  |                                 
     |  |                       +--+----- Anemina_arcaeformis[8] **     
     |  |                          |                                 
     |  |                          +----- Anemina_arcaeformis[9] **     
     |  |                          |                                 
     |  |                          |  +-- Anemina_arcaeformis[1] **     
     |  |                          +--+                              
     |  |                             +-- Anemina_euscaphys **          
     +--+                                                            
        |                             +-- Sinanodonta_angula **         
        |                 +-----------+                              
        |                 |           +-- Sinanodonta_lucida **         
        |  +--------------+                                          
        |  |              |  +----------- Sinanodonta_woodiana[2] **    
        |  |              +--+                                       
        |  |                 |  +-------- Sinanodonta_jourdyi **        
        |  |                 +--+                                    
        |  |                    |  +----- Sinanodonta_elliptica **      
        |  |                    +--+                                 
        |  |                       |  +-- Sinanodonta_woodiana[1] **    
        |  |                       +--+                              
        |  |                          +-- Sinanodonta_lauta **          
        |  |                                                         
        |  |              +-------------- Acuticosta_ovata[1] **        
        |  |              |                                          
        |  |              |  +----------- Acuticosta_chinensis[3] **    
        +--+  +-----------+  |                                       
           |  |           |  +----------- Acuticosta_chinensis[4] **    
           |  |           |  |                                       
           |  |           +--+----------- Acuticosta_chinensis[6] **    
           |  |              |                                       
           |  |              |        +-- Acuticosta_chinensis[1] **    
           |  |              |  +-----+                              
           |  |              |  |     +-- Acuticosta_ovata[2] **        
           |  |              +--+                                    
           |  |                 |  +----- Acuticosta_chinensis[5] **    
           |  |                 |  |                                 
           |  |                 +--+----- Acuticosta_chinensis[7] **    
           |  |                    |                                 
           |  |                    |  +-- Acuticosta_chinensis[2] **    
           +--+                    +--+                              
              |                       +-- Acuticosta_ovata[3] **        
              |                                                      
              |                       +-- Lepidodesma_languilati **     
              |  +--------------------+                              
              |  |                    +-- Cristaria_plicata **          
              |  |                                                   
              |  |           +----------- Lanceolaria_gladiola **       
              |  |           |                                       
              |  |           |  +-------- Lanceolaria_grayii[5] **      
              |  |  +--------+  |                                    
              |  |  |        |  |     +-- Lanceolaria_lanceolata **     
              |  |  |        |  +-----+                              
              |  |  |        |  |     +-- Lanceolaria_triformis **      
              +--+  |        +--+                                    
                 |  |           |  +----- Lanceolaria_grayii[4] **      
                 |  |           |  |                                 
                 |  |           |  |  +-- Lanceolaria_eucylindrica[1] **
                 |  |           |  |  |                              
                 |  |           +--+  +-- Lanceolaria_eucylindrica[2] **
                 |  |              |  |                              
                 |  |              |  +-- Lanceolaria_eucylindrica[3] **
                 |  |              |  |                              
                 |  |              +--+-- Lanceolaria_eucylindrica[4] **
                 +--+                 |                              
                    |                 +-- Lanceolaria_grayii[1] **      
                    |                 |                              
                    |                 +-- Lanceolaria_grayii[2] **      
                    |                 |                              
                    |                 +-- Lanceolaria_grayii[3] **      
                    |                                                
                    |        +----------- Aculamprotula_tortuosa **     
                    |  +-----+                                       
                    |  |     |  +-------- Aculamprotula_tientsinensis **
                    |  |     +--+                                    
                    |  |        |  +----- Aculamprotula_fibrosa **      
                    |  |        +--+                                 
                    |  |           |  +-- Aculamprotula_coreana **      
                    +--+           +--+                              
                       |              +-- Aculamprotula_scripta **      
                       |                                             
                       |           +----- Nodularia_douglasiae[1] **    
                       |  +--------+                                 
                       |  |        |  +-- Nodularia_nuxpersicae **      
                       |  |        +--+                              
                       +--+           +-- Nodularia_douglasiae[2] **    
                          |                                          
                          |  +----------- Tchangsinaia_piscicula **     
                          |  |                                       
                          +--+        +-- Schistodesmus_spinosus **     
                             |  +-----+                              
                             |  |     +-- Schistodesmus_lampreyanus **  
                             +--+                                    
                                |  +----- Cuneopsis_rufescens **        
                                +--+                                 
                                   |  +-- Cuneopsis_heudei **           
                                   +--+                              
                                      +-- Cuneopsis_celtiformis **      

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
69 70 63 4 12 2 1 95.0
                       +-------- Trapezidens_dolichorhynchus **
  +--------------------+                                    
  |                    |  +----- Leoparreysia_tavoyensis **    
  |                    +--+                                 
  |                       |  +-- Coelatura_aegyptiaca **       
  |                       +--+                              
  |                          +-- Indochinella_pugio **         
  |                                                         
  |                       +----- Unio_gibbus **                
  |                       |                                 
  |           +-----------+----- Unio_tumidus **               
  |           |           |                                 
  |           |           |  +-- Unio_crassus **               
  |           |           +--+                              
  |        +--+              +-- Unio_pictorum **              
  |        |  |                                             
  |        |  |  +-------------- Aculamprotula_tortuosa **     
  |        |  |  |                                          
  |        |  +--+           +-- Nodularia_nuxpersicae **      
  |        |     |  +--------+                              
  |        |     |  |        +-- Nodularia_douglasiae **       
  |        |     +--+                                       
  |        |        |  +-------- Schistodesmus_lampreyanus **  
  |        |        +--+                                    
  |        |           |  +----- Tchangsinaia_piscicula **     
  |        |           +--+                                 
  |  +-----+              |  +-- Cuneopsis_heudei **           
  |  |     |              +--+                              
  |  |     |                 +-- Cuneopsis_rufescens **        
--+  |     |                                                
  |  |     |           +-------- Lanceolaria_gladiola **       
  |  |     |  +--------+                                    
  |  |     |  |        |  +----- Lanceolaria_grayii[2] **      
  |  |     |  |        +--+                                 
  |  |     |  |           |  +-- Lanceolaria_lanceolata **     
  |  |     |  |           +--+                              
  |  |     |  |              +-- Lanceolaria_grayii[1] **      
  |  |     |  |                                             
  |  |     +--+     +----------- Cristaria_plicata **          
  |  |        |  +--+                                       
  |  |        |  |  |  +-------- Anemina_arcaeformis **        
  |  |        |  |  +--+                                    
  |  |        |  |     |  +----- Pletholophus_tenuis **        
  |  |        |  |     +--+                                 
  |  |        |  |        |  +-- Sinanodonta_jourdyi **        
  |  |        |  |        +--+                              
  |  |        +--+           +-- Sinanodonta_lucida **         
  |  |           |                                          
  |  |           |        +----- Pseudanodonta_complanata **   
  |  |           |        |                                 
  |  |           |  +-----+  +-- Anodonta_anatina **           
  |  |           |  |     |  |                              
  |  |           |  |     +--+-- Anodonta_cygnea **            
  |  |           |  |        |                              
  |  |           +--+        +-- Anodonta_nuttalliana **       
  |  |              |                                       
  |  |              |        +-- Pyganodon_grandis **          
  +--+              |  +-----+                              
     |              |  |     +-- Simpsonaias_ambigua **        
     |              +--+                                    
     |                 |  +----- Strophitus_undulatus **       
     |                 +--+                                 
     |                    |  +-- Alasmidonta_marginata **      
     |                    +--+                              
     |                       +-- Lasmigona_costata **          
     |                                                      
     |                       +-- Lens_contradens **            
     |                    +--+                              
     |                    |  +-- Lens_comptus **               
     |                 +--+                                 
     |                 |  |  +-- Physunio_modelli **           
     |                 |  +--+                              
     |                 |     +-- unionoid_species           
     |                 |                                    
     |  +--------------+  +----- Hyriopsis_desowitzi **        
     |  |              |  |                                 
     |  |              |  +----- Rectidens_sumatrensis **      
     |  |              |  |                                 
     |  |              |  |  +-- Hyriopsis_khoratensis **      
     |  |              +--+--+                              
     |  |                 |  +-- Hyriopsis_myersiana **        
     |  |                 |                                 
     |  |                 |  +-- Ensidens_ingallsianus **      
     |  |                 |  |                              
     |  |                 +--+-- Ensidens_telus **             
     |  |                    |                              
     |  |                    +-- Ensidens_jaculus **           
     |  |                                                   
     |  |              +-------- Amblema_plicata **            
     +--+           +--+                                    
        |           |  |  +----- Cambarunio_iris **            
        |           |  +--+                                 
        |           |     |  +-- Lampsilis_cardium **          
        |           |     +--+                              
        |  +--------+        +-- Ortmanniana_ligamentina **    
        |  |        |                                       
        |  |        |        +-- Quadrula_quadrula **          
        |  |        |  +-----+                              
        |  |        |  |     +-- Quadrula_verrucosa **         
        |  |        +--+                                    
        |  |           |  +----- Elliptio_complanata **        
        |  |           +--+                                 
        |  |              |  +-- Pleurobema_sintoxia **        
        |  |              +--+                              
        |  |                 +-- Eurynia_dilatata **           
        +--+                                                
           |                 +-- Chamberlainia_somsakpanhai ** 
           |           +-----+                              
           |           |     +-- Sinohyriopsis_cumingii **     
           |           |                                    
           |  +--------+     +-- Lamprotula_caveata **         
           |  |        |  +--+                              
           |  |        |  |  +-- Lamprotula_leaii **           
           |  |        +--+                                 
           |  |           |  +-- Potomida_littoralis **        
           |  |           +--+                              
           |  |              +-- Pronodularia_japanensis **    
           +--+                                             
              |              +-- Sinosolenaia_carinata **      
              |        +-----+                              
              |        |     +-- Sinosolenaia_oleivora **      
              |  +-----+                                    
              |  |     |  +----- Gonidea_angulata **           
              |  |     +--+                                 
              |  |        |  +-- Microcondylaea_bonellii **    
              +--+        +--+                              
                 |           +-- Leguminaia_wheatleyi **       
                 |                                          
                 |  +----------- Parvasolenaia_rivularis **    
                 |  |                                       
                 +--+        +-- Pilsbryoconcha_exilis **      
                    |  +-----+                              
                    |  |     +-- Pilsbryoconcha_acuta **       
                    +--+                                    
                       |  +----- Pseudodon_vagulus **          
                       +--+                                 
                          |  +-- Bineurus_exilis **            
                          +--+                              
                             +-- Nyeinchanconcha_nyeinchani ** 

Jeratthitikul et al., 2022 [Publ 4260], fig. S1, 1

iT tT bi riS oT roT roS iD
66 66 63 3 10 10 1 88.0
                    +-------------------- Cumberlandia_monodonta      
                    |                                                 
  +-----------------+  +----------------- Gibbosula_laosensis         
  |                 |  |                                              
  |                 |  |              +-- Pseudunio_auricularius      
  |                 +--+  +-----------+                               
  |                    |  |           +-- Pseudunio_marocanus         
  |                    |  |                                           
  |                    +--+           +-- Margaritifera_margaritifera 
  |                       |  +--------+                               
  |                       |  |        +-- Margaritifera_dahurica      
  |                       +--+                                        
  |                          |  +-------- Margaritifera_falcata       
  |                          +--+                                     
  |                             |  +----- Margaritifera_middendorffi  
  |                             +--+                                  
  |                                |  +-- Margaritifera_laevis        
  |                                +--+                               
  |                                   +-- Margaritifera_marrianae     
  |                                                                   
  |                                   +-- Indopseudodon_indawgyiensis ** 
  |              +--------------------+                               
  |              |                    +-- Indopseudodon_kayinensis **    
  |              |                                                    
--+              |                    +-- Nyeinchanconcha_nyeinchani **  
  |              |              +-----+                               
  |  +-----------+              |     +-- Sundadontina_cumingii **       
  |  |           |  +-----------+                                     
  |  |           |  |           |  +----- Thaiconcha_callifera **        
  |  |           |  |           +--+                                  
  |  |           |  |              |  +-- Bineurus_panhai **             
  |  |           |  |              +--+                               
  |  |           +--+                 +-- Bineurus_mouhotii **           
  |  |              |                                                 
  |  |              |                 +-- Namkongnaia_inkhavilayi **     
  |  |              |  +--------------+                               
  |  |              |  |              +-- Pseudodon_vondembuschianus **  
  |  |              |  |                                              
  |  |              +--+              +-- Pilsbryoconcha_exilis **       
  |  |                 |  +-----------+                               
  |  |                 |  |           +-- Pilsbryoconcha_schomburgki **  
  |  |                 |  |                                           
  |  |                 +--+           +-- Pilsbryoconcha_carinifera **   
  |  |                    |  +--------+                               
  |  |                    |  |        +-- Pilsbryoconcha_linguaeformis **
  |  |                    +--+                                        
  +--+                       |  +-------- Pilsbryoconcha_hoikaab **      
     |                       +--+                                     
     |                          |  +----- Pilsbryoconcha_mekongiana **   
     |                          +--+                                  
     |                             |  +-- Pilsbryoconcha_acuta **        
     |                             +--+                               
     |                                +-- Pilsbryoconcha_kittitati **    
     |                                                                
     |                             +----- Lens_contradens **             
     |                          +--+                                  
     |                          |  |  +-- Trapezoideus_foliaceus **      
     |                          |  +--+                               
     |  +-----------------------+     +-- Physunio_superbus **           
     |  |                       |                                     
     |  |                       |     +-- Ensidens_ingallsianus **       
     |  |                       |     |                               
     |  |                       +-----+-- Hyriopsis_bialata **           
     |  |                             |                               
     |  |                             +-- Rectidens_sumatrensis **       
     |  |                                                             
     |  |                          +----- Sinosolenaia_carinata **       
     +--+  +-----------------------+                                  
        |  |                       |  +-- Leguminaia_wheatleyi **        
        |  |                       +--+                               
        |  |                          +-- Gonidea_angulata **            
        |  |                                                          
        |  |                          +-- Chamberlainia_somsakpanhai **  
        |  |                    +-----+                               
        |  |                    |     +-- Sinohyriopsis_cumingii **      
        |  |  +-----------------+                                     
        |  |  |                 |  +----- Lamprotula_leaii **            
        +--+  |                 +--+                                  
           |  |                    |  +-- Potomida_littoralis **         
           |  |                    +--+                               
           |  |                       +-- Pronodularia_japanensis **     
           |  |                                                       
           |  |                 +-------- Amblema_plicata **             
           |  |              +--+                                     
           |  |              |  |  +----- Ortmanniana_ligamentina **     
           |  |              |  +--+                                  
           |  |              |     |  +-- Cambarunio_iris **             
           |  |              |     +--+                               
           +--+  +-----------+        +-- Lampsilis_cardium **           
              |  |           |                                        
              |  |           |        +-- Quadrula_quadrula **           
              |  |           |  +-----+                               
              |  |           |  |     +-- Quadrula_verrucosa **          
              |  |           +--+                                     
              |  |              |  +----- Elliptio_complanata **         
              |  |              +--+                                  
              |  |                 |  +-- Pleurobema_sintoxia **         
              |  |                 +--+                               
              |  |                    +-- Eurynia_dilatata **            
              |  |                                                    
              |  |                 +----- Unio_tumidus **                
              +--+     +-----------+                                  
                 |     |           |  +-- Unio_pictorum **               
                 |     |           +--+                               
                 |  +--+              +-- Unio_crassus **                
                 |  |  |                                              
                 |  |  |  +-------------- Aculamprotula_tortuosa **      
                 |  |  |  |                                           
                 |  |  +--+           +-- Nodularia_douglasiae **        
                 |  |     |  +--------+                               
                 |  |     |  |        +-- Nodularia_nuxpersicae **       
                 |  |     +--+                                        
                 |  |        |  +-------- Schistodesmus_lampreyanus **   
                 |  |        +--+                                     
                 |  |           |  +----- Cuneopsis_heudei **            
                 +--+           +--+                                  
                    |              |  +-- Tchangsinaia_piscicula **      
                    |              +--+                               
                    |                 +-- Cuneopsis_rufescens **         
                    |                                                 
                    |                 +-- Lanceolaria_grayii **          
                    |  +--------------+                               
                    |  |              +-- Lanceolaria_gladiola **        
                    |  |                                              
                    |  |     +----------- Cristaria_plicata **           
                    |  |  +--+                                        
                    +--+  |  |  +-------- Pletholophus_tenuis **         
                       |  |  +--+                                     
                       |  |     |  +----- Sinanodonta_lucida **          
                       |  |     +--+                                  
                       |  |        |  +-- Sinanodonta_lauta **           
                       |  |        +--+                               
                       +--+           +-- Sinanodonta_jourdyi **         
                          |                                           
                          |     +-------- Pseudanodonta_complanata **    
                          |  +--+                                     
                          |  |  |  +----- Anodonta_nuttalliana **        
                          |  |  +--+                                  
                          |  |     |  +-- Anodonta_anatina **            
                          |  |     +--+                               
                          +--+        +-- Anodonta_cygnea **             
                             |                                        
                             |        +-- Pyganodon_grandis **           
                             |  +-----+                               
                             |  |     +-- Simpsonaias_ambigua **         
                             +--+                                     
                                |  +----- Strophitus_undulatus **        
                                +--+                                  
                                   |  +-- Alasmidonta_marginata **       
                                   +--+                               
                                      +-- Lasmigona_costata **           

Jeratthitikul et al., 2022 [Publ 4260], fig. S2, 2

iT tT bi riS oT roT roS iD
66 66 64 3 10 10 1 86.0
                       +----------------- Cumberlandia_monodonta      
                       |                                              
                       |           +----- Gibbosula_laosensis         
  +--------------------+  +--------+                                  
  |                    |  |        |  +-- Pseudunio_auricularius      
  |                    |  |        +--+                               
  |                    |  |           +-- Pseudunio_marocanus         
  |                    +--+                                           
  |                       |           +-- Margaritifera_dahurica      
  |                       |  +--------+                               
  |                       |  |        +-- Margaritifera_margaritifera 
  |                       +--+                                        
  |                          |  +-------- Margaritifera_falcata       
  |                          +--+                                     
  |                             |  +----- Margaritifera_marrianae     
  |                             +--+                                  
  |                                |  +-- Margaritifera_middendorffi  
  |                                +--+                               
  |                                   +-- Margaritifera_laevis        
  |                                                                   
  |                                   +-- Indopseudodon_indawgyiensis ** 
  |              +--------------------+                               
  |              |                    +-- Indopseudodon_kayinensis **    
  |              |                                                    
--+              |                    +-- Nyeinchanconcha_nyeinchani **  
  |              |              +-----+                               
  |  +-----------+              |     +-- Sundadontina_cumingii **       
  |  |           |  +-----------+                                     
  |  |           |  |           |  +----- Thaiconcha_callifera **        
  |  |           |  |           +--+                                  
  |  |           |  |              |  +-- Bineurus_panhai **             
  |  |           |  |              +--+                               
  |  |           +--+                 +-- Bineurus_mouhotii **           
  |  |              |                                                 
  |  |              |                 +-- Namkongnaia_inkhavilayi **     
  |  |              |  +--------------+                               
  |  |              |  |              +-- Pseudodon_vondembuschianus **  
  |  |              |  |                                              
  |  |              +--+              +-- Pilsbryoconcha_exilis **       
  |  |                 |  +-----------+                               
  |  |                 |  |           +-- Pilsbryoconcha_schomburgki **  
  |  |                 |  |                                           
  |  |                 +--+           +-- Pilsbryoconcha_carinifera **   
  |  |                    |  +--------+                               
  |  |                    |  |        +-- Pilsbryoconcha_linguaeformis **
  |  |                    +--+                                        
  +--+                       |  +-------- Pilsbryoconcha_hoikaab **      
     |                       +--+                                     
     |                          |  +----- Pilsbryoconcha_mekongiana **   
     |                          +--+                                  
     |                             |  +-- Pilsbryoconcha_acuta **        
     |                             +--+                               
     |                                +-- Pilsbryoconcha_kittitati **    
     |                                                                
     |                             +----- Ensidens_ingallsianus **       
     |                          +--+                                  
     |                          |  |  +-- Rectidens_sumatrensis **       
     |                          |  +--+                               
     |  +-----------------------+     +-- Hyriopsis_bialata **           
     |  |                       |                                     
     |  |                       |  +----- Lens_contradens **             
     |  |                       +--+                                  
     |  |                          |  +-- Trapezoideus_foliaceus **      
     |  |                          +--+                               
     |  |                             +-- Physunio_superbus **           
     |  |                                                             
     |  |                          +----- Sinosolenaia_carinata **       
     +--+  +-----------------------+                                  
        |  |                       |  +-- Gonidea_angulata **            
        |  |                       +--+                               
        |  |                          +-- Leguminaia_wheatleyi **        
        |  |                                                          
        |  |                          +-- Chamberlainia_somsakpanhai **  
        |  |                    +-----+                               
        |  |                    |     +-- Sinohyriopsis_cumingii **      
        |  |  +-----------------+                                     
        |  |  |                 |  +----- Potomida_littoralis **         
        +--+  |                 +--+                                  
           |  |                    |  +-- Lamprotula_leaii **            
           |  |                    +--+                               
           |  |                       +-- Pronodularia_japanensis **     
           |  |                                                       
           |  |                 +-------- Amblema_plicata **             
           |  |              +--+                                     
           |  |              |  |  +----- Ortmanniana_ligamentina **     
           |  |              |  +--+                                  
           |  |              |     |  +-- Lampsilis_cardium **           
           |  |              |     +--+                               
           +--+  +-----------+        +-- Cambarunio_iris **             
              |  |           |                                        
              |  |           |        +-- Quadrula_quadrula **           
              |  |           |  +-----+                               
              |  |           |  |     +-- Quadrula_verrucosa **          
              |  |           +--+                                     
              |  |              |  +----- Pleurobema_sintoxia **         
              |  |              +--+                                  
              |  |                 |  +-- Eurynia_dilatata **            
              |  |                 +--+                               
              |  |                    +-- Elliptio_complanata **         
              |  |                                                    
              |  |                 +----- Unio_tumidus **                
              +--+     +-----------+                                  
                 |     |           |  +-- Unio_pictorum **               
                 |     |           +--+                               
                 |  +--+              +-- Unio_crassus **                
                 |  |  |                                              
                 |  |  |  +-------------- Aculamprotula_tortuosa **      
                 |  |  |  |                                           
                 |  |  +--+           +-- Nodularia_douglasiae **        
                 |  |     |  +--------+                               
                 |  |     |  |        +-- Nodularia_nuxpersicae **       
                 |  |     +--+                                        
                 |  |        |  +-------- Schistodesmus_lampreyanus **   
                 |  |        +--+                                     
                 |  |           |  +----- Cuneopsis_heudei **            
                 +--+           +--+                                  
                    |              |  +-- Tchangsinaia_piscicula **      
                    |              +--+                               
                    |                 +-- Cuneopsis_rufescens **         
                    |                                                 
                    |                 +-- Lanceolaria_grayii **          
                    |  +--------------+                               
                    |  |              +-- Lanceolaria_gladiola **        
                    |  |                                              
                    |  |     +----------- Cristaria_plicata **           
                    |  |  +--+                                        
                    +--+  |  |  +-------- Pletholophus_tenuis **         
                       |  |  +--+                                     
                       |  |     |  +----- Sinanodonta_lucida **          
                       |  |     +--+                                  
                       |  |        |  +-- Sinanodonta_jourdyi **         
                       |  |        +--+                               
                       +--+           +-- Sinanodonta_lauta **           
                          |                                           
                          |     +-------- Pseudanodonta_complanata **    
                          |  +--+                                     
                          |  |  |  +----- Anodonta_anatina **            
                          |  |  +--+                                  
                          |  |     |  +-- Anodonta_nuttalliana **        
                          |  |     +--+                               
                          +--+        +-- Anodonta_cygnea **             
                             |                                        
                             |        +-- Alasmidonta_marginata **       
                             |  +-----+                               
                             |  |     +-- Lasmigona_costata **           
                             +--+                                     
                                |  +----- Pyganodon_grandis **           
                                +--+                                  
                                   |  +-- Strophitus_undulatus **        
                                   +--+                               
                                      +-- Simpsonaias_ambigua **         

Wu et al., 2022 [Publ 4238], fig. 3

iT tT bi riS oT roT roS iD
65 65 63 3 9 6 1 100.0
                             +-------------- Gibbosula_rochechouartii    
  +--------------------------+                                           
  |                          |  +----------- Cumberlandia_monodonta      
  |                          +--+                                        
  |                             |  +-------- Pseudunio_marocanus         
  |                             +--+                                     
  |                                |  +----- Margaritifera_falcata       
  |                                +--+                                  
  |                                   |  +-- Margaritifera_dahurica      
  |                                   +--+                               
  |                                      +-- Margaritifera_margaritifera 
  |                                                                      
  |                                      +-- Uniomerus_tetralasmus **       
  |           +--------------------------+                               
  |           |                          +-- Quadrula_quadrula **           
  |           |                                                          
  |  +--------+                          +-- Pleurobema_oviforme **         
  |  |        |  +-----------------------+                               
  |  |        |  |                       +-- Elliptio_complanata **         
  |  |        +--+                                                       
--+  |           |  +----------------------- Amblema_plicata **             
  |  |           |  |                                                    
  |  |           +--+  +-------------------- Popenaias_popeii **            
  |  |              |  |                                                 
  |  |              +--+  +----------------- Toxolasma_parvum **            
  |  |                 |  |                                              
  |  |                 +--+              +-- Potamilus_alatus **            
  |  |                    |  +-----------+                               
  |  |                    |  |           +-- Potamilus_leptodon **          
  |  |                    +--+                                           
  |  |                       |  +----------- Leaunio_lienosus **            
  |  |                       |  |                                        
  |  |                       +--+        +-- Lampsilis_cardium **           
  |  |                          |  +-----+                               
  |  |                          |  |     +-- Lampsilis_ornata **            
  |  |                          +--+                                     
  |  |                             |  +----- Venustaconcha_ellipsiformis ** 
  |  |                             +--+                                  
  |  |                                |  +-- Lampsilis_siliquoidea **       
  |  |                                +--+                               
  +--+                                   +-- Lampsilis_powellii **          
     |                                                                   
     |                                +----- Chamberlainia_somsakpanhai **  
     |                    +-----------+                                  
     |                    |           |  +-- Sinohyriopsis_schlegelii **    
     |                    |           +--+                               
     |                    |              +-- Sinohyriopsis_cumingii **      
     |                    |                                              
     |  +-----------------+        +-------- Potomida_littoralis **         
     |  |                 |  +-----+                                     
     |  |                 |  |     |  +----- Pronodularia_japanensis **     
     |  |                 |  |     +--+                                  
     |  |                 |  |        |  +-- Lamprotula_caveata **          
     |  |                 |  |        +--+                               
     |  |                 +--+           +-- Lamprotula_leaii **            
     |  |                    |                                           
     |  |                    |           +-- Pilsbryoconcha_exilis **       
     |  |                    |  +--------+                               
     |  |                    |  |        +-- Pseudodon_vondembuschianus **  
     |  |                    |  |                                        
     |  |                    +--+        +-- Ptychorhynchus_pfisteri **     
     |  |                       |  +-----+                               
     |  |                       |  |     +-- Parvasolenaia_rivularis **     
     |  |                       +--+                                     
     |  |                          |  +----- Microcondylaea_bonellii **     
     +--+                          +--+                                  
        |                             |  +-- Sinosolenaia_carinata **       
        |                             +--+                               
        |                                +-- Sinosolenaia_recognita **      
        |                                                                
        |  +-------------------------------- Lepidodesma_languilati **      
        |  |                                                             
        |  |                       +-------- Acuticosta_chinensis **        
        |  |              +--------+                                     
        |  |              |        |  +----- Lanceolaria_gladiola **        
        |  |              |        +--+                                  
        |  |              |           |  +-- Lanceolaria_grayii **          
        |  |              |           +--+                               
        |  |              |              +-- Lanceolaria_lanceolata **      
        |  |              |                                              
        |  |  +-----------+        +-------- Cristaria_plicata **           
        |  |  |           |        |                                     
        +--+  |           |  +-----+     +-- Sinanodonta_woodiana **        
           |  |           |  |     |  +--+                               
           |  |           |  |     |  |  +-- Sinanodonta_lucida **          
           |  |           |  |     +--+                                  
           |  |           |  |        |  +-- Anemina_arcaeformis **         
           |  |           +--+        +--+                               
           |  |              |           +-- Anemina_euscaphys **           
           |  |              |                                           
           |  |              |           +-- Anodonta_cygnea **             
           |  |              |  +--------+                               
           |  |              |  |        +-- Anodonta_anatina **            
           |  |              +--+                                        
           |  |                 |  +-------- Platynaias_compressa **        
           +--+                 +--+                                     
              |                    |  +----- Pyganodon_grandis **           
              |                    +--+                                  
              |                       |  +-- Utterbackia_imbecillis **      
              |                       +--+                               
              |                          +-- Utterbackia_peninsularis **    
              |                                                          
              |                    +-------- Aculamprotula_tortuosa **      
              |                    |                                     
              |  +-----------------+     +-- Aculamprotula_tientsinensis ** 
              |  |                 |  +--+                               
              |  |                 |  |  +-- Aculamprotula_coreana **       
              |  |                 +--+                                  
              |  |                    |  +-- Aculamprotula_polysticta **    
              |  |                    +--+                               
              |  |                       +-- Aculamprotula_scripta **       
              +--+                                                       
                 |                 +-------- Unio_tumidus **                
                 |  +--------------+                                     
                 |  |              |  +----- Unio_crassus **                
                 |  |              +--+                                  
                 |  |                 |  +-- Unio_pictorum **               
                 |  |                 +--+                               
                 +--+                    +-- Unio_delphinus **              
                    |                                                    
                    |                    +-- Pseudocuneopsis_capitata **    
                    |  +-----------------+                               
                    |  |                 +-- Pseudocuneopsis_sichuanensis **
                    |  |                                                 
                    +--+                 +-- Nodularia_douglasiae **        
                       |  +--------------+                               
                       |  |              +-- Nodularia_breviconcha **       
                       |  |                                              
                       +--+              +-- Schistodesmus_lampreyanus **   
                          |  +-----------+                               
                          |  |           +-- Schistodesmus_spinosus **      
                          +--+                                           
                             |  +----------- Tchangsinaia_piscicula **      
                             +--+                                        
                                |  +-------- Cuneopsis_rufescens **         
                                +--+                                     
                                   |  +----- Cuneopsis_celtiformis **       
                                   +--+                                  
                                      |  +-- Cuneopsis_heudei **            
                                      +--+                               
                                         +-- Cuneopsis_demangei **          

Bolotov et al., 2022 [Publ 4252], fig. 2

iT tT bi riS oT roT roS iD
61 61 55 2 0 0 0 103.0
                                               +-- Pseudodon_lenyanensis **        
                                               |                                
                                               +-- Nyeinchanconcha_nyeinchani **   
  +--------------------------------------------+                                
  |                                            +-- Indopseudodon_kayinensis **     
  |                                            |                                
  |                                            +-- Sundadontina_tanintharyiensis **
  |                                                                             
  |                                   +----------- Trapezidens_mogaungensis **     
  |                                   |                                         
  |                             +-----+  +-------- Trapezidens_exolescens **       
  |                             |     |  |                                      
  |                             |     +--+  +----- Trapezidens_scutum **           
  |                             |        |  |                                   
  |                             |        +--+  +-- Trapezidens_angustior **        
  |                             |           |  |                                
  |  +--------------------------+           +--+-- Trapezidens_dolichorhynchus **  
  |  |                          |              |                                
--+  |                          |              +-- Trapezidens_yeti **             
  |  |                          |                                               
  |  |                          |  +-------------- Lamellidens_marginalis **       
  |  |                          |  |                                            
  |  |                          |  |  +----------- Lamellidens_ferrugineus **      
  |  |                          +--+  |                                         
  |  |                             |  |  +-------- Lamellidens_generosus **        
  |  |                             |  +--+                                      
  |  |                             |  |  |  +----- Lamellidens_mainwaringi **      
  |  |                             +--+  +--+                                   
  |  |                                |     |  +-- Lamellidens_lamellatus **       
  |  |                                |     +--+                                
  |  |                                |        +-- Lamellidens_jenkinsianus **     
  |  |                                |                                         
  |  |                                |  +-------- Lamellidens_nongyangensis **    
  +--+                                +--+                                      
     |                                   |  +----- Lamellidens_savadiensis **      
     |                                   +--+                                   
     |                                      |  +-- Lamellidens_corrianus **        
     |                                      +--+                                
     |                                         +-- Lamellidens_chindwinensis **    
     |                                                                          
     |                                +----------- Parreysia_keralaensis **        
     |  +-----------------------------+                                         
     |  |                             |  +-------- Parreysia_corrugata **          
     |  |                             +--+                                      
     |  |                                |  +----- Parreysia_rakhinensis **        
     |  |                                +--+                                   
     |  |                                   |  +-- Parreysia_nagpoorensis **       
     |  |                                   +--+                                
     |  |                                      +-- Parreysia_plagiosoma **         
     |  |                                                                       
     |  |                                      +-- Leoparreysia_burmana **         
     +--+                                +-----+                                
        |                                |     +-- Leoparreysia_whitteni **        
        |  +-----------------------------+                                      
        |  |                             |  +----- Leoparreysia_olivacea **        
        |  |                             +--+                                   
        |  |                                |  +-- Leoparreysia_canefrii **        
        |  |                                +--+                                
        |  |                                   +-- Leoparreysia_tavoyensis **      
        |  |                                                                    
        |  |                             +-------- Nitia_teretiuscula **           
        +--+  +--------------------------+                                      
           |  |                          |  +----- Coelatura_aegyptiaca **         
           |  |                          +--+                                   
           |  |                             |  +-- Nyassunio_nyassaensis **        
           |  |                             +--+                                
           |  |                                +-- Coelatura_kunenensis **         
           |  |                                                                 
           |  |                 +----------------- Indonaia_cylindrica **          
           |  |                 |                                               
           +--+  +--------------+  +-------------- Indonaia_gratiosa **            
              |  |              |  |                                            
              |  |              +--+  +----------- Indonaia_andersoniana **        
              |  |                 |  |                                         
              |  |                 +--+        +-- Indonaia_caerulea **            
              |  |                    |  +-----+                                
              |  |                    |  |     +-- Indonaia_subclathrata **        
              |  |                    +--+                                      
              |  |                       |  +----- Indonaia_rugosa **              
              +--+                       +--+                                   
                 |                          |  +-- Indonaia_bonneaudii **          
                 |                          +--+                                
                 |                             +-- Indonaia_shurtleffiana **       
                 |                                                              
                 |  +----------------------------- Indochinella_pugio **           
                 |  |                                                           
                 |  |                 +----------- Radiatula_crispisulcata **      
                 |  |                 |                                         
                 |  |                 |     +----- Radiatula_chaudhurii **         
                 +--+  +--------------+  +--+                                   
                    |  |              |  |  |  +-- Radiatula_myitkyinae **         
                    |  |              |  |  +--+                                
                    |  |              +--+     +-- Radiatula_kachinensis **        
                    |  |                 |                                      
                    |  |                 |  +----- Radiatula_chindwinensis **      
                    +--+                 +--+                                   
                       |                    |  +-- Radiatula_myitthanensis **      
                       |                    +--+                                
                       |                       +-- Radiatula_mouhoti **            
                       |                                                        
                       |  +----------------------- Scabiellus_songkramensis **     
                       |  |                                                     
                       +--+  +-------------------- Harmandia_somboriensis **       
                          |  |                                                  
                          +--+  +----------------- Unionetta_fabagina **           
                             |  |                                               
                             +--+              +-- Scabies_crispata **             
                                |  +-----------+                                
                                |  |           +-- Scabies_mandarinus **           
                                +--+                                            
                                   |  +----------- Scabies_nucleus **              
                                   |  |                                         
                                   +--+        +-- Scabies_humilis **              
                                      |  +-----+                                
                                      |  |     +-- Scabies_pilata **               
                                      +--+                                      
                                         |  +----- Scabies_phaselus **             
                                         +--+                                   
                                            |  +-- Scabies_anceps **               
                                            +--+                                
                                               +-- Scabies_scobinatus **           

Bolotov et al., 2022 [Publ 4252], fig. S1

iT tT bi riS oT roT roS iD
61 61 49 2 0 0 0 103.0
                                                  +-- Pseudodon_lenyanensis **        
                                                  |                                
                                                  +-- Nyeinchanconcha_nyeinchani **   
  +-----------------------------------------------+                                
  |                                               +-- Indopseudodon_kayinensis **     
  |                                               |                                
  |                                               +-- Sundadontina_tanintharyiensis **
  |                                                                                
  |                                               +-- Parreysia_keralaensis **        
  |                                               |                                
  |                                               +-- Parreysia_corrugata **          
  |                                               |                                
  |  +--------------------------------------------+-- Parreysia_plagiosoma **         
  |  |                                            |                                
  |  |                                            +-- Parreysia_rakhinensis **        
--+  |                                            |                                
  |  |                                            +-- Parreysia_nagpoorensis **       
  |  |                                                                             
  |  |                                +-------------- Trapezidens_mogaungensis **     
  |  |                             +--+                                            
  |  |                             |  |  +----------- Trapezidens_scutum **           
  |  |                             |  +--+                                         
  |  |                             |     |  +-------- Trapezidens_exolescens **       
  |  |                             |     +--+                                      
  |  |                             |        |  +----- Trapezidens_yeti **             
  |  |                             |        +--+                                   
  |  |                             |           |  +-- Trapezidens_dolichorhynchus **  
  |  |  +--------------------------+           +--+                                
  +--+  |                          |              +-- Trapezidens_angustior **        
     |  |                          |                                               
     |  |                          |        +-------- Lamellidens_savadiensis **      
     |  |                          |  +-----+                                      
     |  |                          |  |     |  +----- Lamellidens_chindwinensis **    
     |  |                          |  |     +--+                                   
     |  |                          |  |        |  +-- Lamellidens_corrianus **        
     |  |                          |  |        +--+                                
     |  |                          +--+           +-- Lamellidens_nongyangensis **    
     |  |                             |                                            
     |  |                             |           +-- Lamellidens_marginalis **       
     |  |                             |  +--------+                                
     |  |                             |  |        +-- Lamellidens_ferrugineus **      
     |  |                             +--+                                         
     |  |                                |  +-------- Lamellidens_generosus **        
     |  |                                +--+                                      
     |  |                                   |  +----- Lamellidens_mainwaringi **      
     +--+                                   +--+                                   
        |                                      |  +-- Lamellidens_jenkinsianus **     
        |                                      +--+                                
        |                                         +-- Lamellidens_lamellatus **       
        |                                                                          
        |                                         +-- Leoparreysia_burmana **         
        |                                         |                                
        |                                         +-- Leoparreysia_canefrii **        
        |                                         |                                
        |  +--------------------------------------+-- Leoparreysia_olivacea **        
        |  |                                      |                                
        |  |                                      +-- Leoparreysia_tavoyensis **      
        |  |                                      |                                
        |  |                                      +-- Leoparreysia_whitteni **        
        |  |                                                                       
        |  |                                      +-- Coelatura_aegyptiaca **         
        |  |                                      |                                
        |  |                                      +-- Coelatura_kunenensis **         
        +--+  +-----------------------------------+                                
           |  |                                   +-- Nitia_teretiuscula **           
           |  |                                   |                                
           |  |                                   +-- Nyassunio_nyassaensis **        
           |  |                                                                    
           |  |                                   +-- Indonaia_cylindrica **          
           |  |                          +--------+                                
           |  |                          |        +-- Indonaia_gratiosa **            
           |  |                          |                                         
           +--+  +-----------------------+     +----- Indonaia_shurtleffiana **       
              |  |                       |  +--+                                   
              |  |                       |  |  |  +-- Indonaia_bonneaudii **          
              |  |                       |  |  +--+                                
              |  |                       +--+     +-- Indonaia_rugosa **              
              |  |                          |                                      
              |  |                          |  +----- Indonaia_andersoniana **        
              |  |                          +--+                                   
              |  |                             |  +-- Indonaia_caerulea **            
              +--+                             +--+                                
                 |                                +-- Indonaia_subclathrata **        
                 |                                                                 
                 |  +-------------------------------- Indochinella_pugio **           
                 |  |                                                              
                 |  |                          +----- Radiatula_chaudhurii **         
                 |  |                    +-----+                                   
                 |  |                    |     |  +-- Radiatula_kachinensis **        
                 |  |                    |     +--+                                
                 +--+  +-----------------+        +-- Radiatula_myitkyinae **         
                    |  |                 |                                         
                    |  |                 |  +-------- Radiatula_crispisulcata **      
                    |  |                 +--+                                      
                    |  |                    |  +----- Radiatula_mouhoti **            
                    |  |                    +--+                                   
                    +--+                       |  +-- Radiatula_myitthanensis **      
                       |                       +--+                                
                       |                          +-- Radiatula_chindwinensis **      
                       |                                                           
                       |  +-------------------------- Scabiellus_songkramensis **     
                       |  |                                                        
                       +--+  +----------------------- Harmandia_somboriensis **       
                          |  |                                                     
                          +--+  +-------------------- Unionetta_fabagina **           
                             |  |                                                  
                             +--+  +----------------- Scabies_crispata **             
                                |  |                                               
                                +--+              +-- Scabies_humilis **              
                                   |  +-----------+                                
                                   |  |           +-- Scabies_mandarinus **           
                                   +--+                                            
                                      |  +----------- Scabies_nucleus **              
                                      +--+                                         
                                         |  +-------- Scabies_pilata **               
                                         +--+                                      
                                            |  +----- Scabies_anceps **               
                                            +--+                                   
                                               |  +-- Scabies_phaselus **             
                                               +--+                                
                                                  +-- Scabies_scobinatus **           

Bolotov et al., 2022 [Publ 4239], fig. 2

iT tT bi riS oT roT roS iD
58 58 54 2 0 0 0 100.0
                                                  +-- Pseudodon_lenyanensis **        
                                                  |                                
                                                  +-- Nyeinchanconcha_nyeinchani **   
  +-----------------------------------------------+                                
  |                                               +-- Indopseudodon_kayinensis **     
  |                                               |                                
  |                                               +-- Sundadontina_tanintharyiensis **
  |                                                                                
  |                                               +-- Trapezidens_exolescens **       
  |                                         +-----+                                
  |                                         |     +-- Trapezidens_scutum **           
  |                             +-----------+                                      
  |                             |           |  +----- Trapezidens_angustior **        
  |                             |           +--+                                   
  |                             |              |  +-- Trapezidens_dolichorhynchus **  
  |  +--------------------------+              +--+                                
  |  |                          |                 +-- Trapezidens_yeti **             
  |  |                          |                                                  
--+  |                          |  +----------------- Lamellidens_corrianus **        
  |  |                          |  |                                               
  |  |                          +--+  +-------------- Lamellidens_nongyangensis **    
  |  |                             |  |                                            
  |  |                             +--+  +----------- Lamellidens_savadiensis **      
  |  |                                |  |                                         
  |  |                                |  |        +-- Lamellidens_ferrugineus **      
  |  |                                +--+  +-----+                                
  |  |                                   |  |     +-- Lamellidens_marginalis **       
  |  |                                   |  |                                      
  |  |                                   +--+     +-- Lamellidens_lamellatus **       
  |  |                                      |  +--+                                
  |  |                                      |  |  +-- Lamellidens_jenkinsianus **     
  |  |                                      +--+                                   
  |  |                                         |  +-- Lamellidens_generosus **        
  |  |                                         +--+                                
  +--+                                            +-- Lamellidens_mainwaringi **      
     |                                                                             
     |                                   +----------- Parreysia_keralaensis **        
     |                                +--+                                         
     |                                |  |  +-------- Parreysia_plagiosoma **         
     |                                |  +--+                                      
     |                                |     |  +----- Parreysia_nagpoorensis **       
     |                                |     +--+                                   
     |                                |        |  +-- Parreysia_corrugata **          
     |  +-----------------------------+        +--+                                
     |  |                             |           +-- Parreysia_rakhinensis **        
     |  |                             |                                            
     |  |                             |           +-- Leoparreysia_burmana **         
     |  |                             |     +-----+                                
     |  |                             |     |     +-- Leoparreysia_whitteni **        
     |  |                             +-----+                                      
     |  |                                   |  +----- Leoparreysia_olivacea **        
     |  |                                   +--+                                   
     |  |                                      |  +-- Leoparreysia_canefrii **        
     |  |                                      +--+                                
     +--+                                         +-- Leoparreysia_tavoyensis **      
        |                                                                          
        |                                   +-------- Nitia_teretiuscula **           
        |  +--------------------------------+                                      
        |  |                                |  +----- Coelatura_aegyptiaca **         
        |  |                                +--+                                   
        |  |                                   |  +-- Coelatura_kunenensis **         
        |  |                                   +--+                                
        |  |                                      +-- Nyassunio_nyassaensis **        
        |  |                                                                       
        |  |                                      +-- Indonaia_cylindrica **          
        |  |                             +--------+                                
        +--+                             |        +-- Indonaia_gratiosa **            
           |                             |                                         
           |  +--------------------------+     +----- Indonaia_andersoniana **        
           |  |                          |  +--+                                   
           |  |                          |  |  |  +-- Indonaia_caerulea **            
           |  |                          |  |  +--+                                
           |  |                          +--+     +-- Indonaia_subclathrata **        
           |  |                             |                                      
           |  |                             |  +----- Indonaia_shurtleffiana **       
           |  |                             +--+                                   
           +--+                                |  +-- Indonaia_bonneaudii **          
              |                                +--+                                
              |                                   +-- Indonaia_rugosa **              
              |                                                                    
              |  +----------------------------------- Indochinella_pugio **           
              |  |                                                                 
              |  |                       +----------- Radiatula_crispisulcata **      
              |  |                       |                                         
              +--+  +--------------------+        +-- Radiatula_chaudhurii **         
                 |  |                    |  +-----+                                
                 |  |                    |  |     +-- Radiatula_myitkyinae **         
                 |  |                    +--+                                      
                 |  |                       |  +----- Radiatula_mouhoti **            
                 +--+                       +--+                                   
                    |                          |  +-- Radiatula_chindwinensis **      
                    |                          +--+                                
                    |                             +-- Radiatula_myitthanensis **      
                    |                                                              
                    |  +----------------------------- Scabiellus_songkramensis **     
                    +--+                                                           
                       |  +-------------------------- Harmandia_somboriensis **       
                       +--+                                                        
                          |  +----------------------- Unionetta_fabagina **           
                          +--+                                                     
                             |  +-------------------- Scabies_crispata **             
                             +--+                                                  
                                |  +----------------- Scabies_mandarinus **           
                                +--+                                               
                                   |  +-------------- Scabies_nucleus **              
                                   +--+                                            
                                      |  +----------- Scabies_pilata **               
                                      +--+                                         
                                         |  +-------- Scabies_humilis **              
                                         +--+                                      
                                            |  +----- Scabies_anceps **               
                                            +--+                                   
                                               |  +-- Scabies_phaselus **             
                                               +--+                                
                                                  +-- Scabies_scobinatus **           

Bolotov et al., 2022 [Publ 4239], fig. S1

iT tT bi riS oT roT roS iD
58 58 52 2 0 0 0 100.0
                                               +-- Pseudodon_lenyanensis **        
                                               |                                
                                               +-- Nyeinchanconcha_nyeinchani **   
  +--------------------------------------------+                                
  |                                            +-- Indopseudodon_kayinensis **     
  |                                            |                                
  |                                            +-- Sundadontina_tanintharyiensis **
  |                                                                             
  |                                      +-------- Trapezidens_exolescens **       
  |                                      |                                      
  |                             +--------+  +----- Trapezidens_scutum **           
  |                             |        |  |                                   
  |                             |        +--+  +-- Trapezidens_angustior **        
  |                             |           |  |                                
  |                             |           +--+-- Trapezidens_dolichorhynchus **  
  |                             |              |                                
  |  +--------------------------+              +-- Trapezidens_yeti **             
--+  |                          |                                               
  |  |                          |           +----- Lamellidens_savadiensis **      
  |  |                          |  +--------+                                   
  |  |                          |  |        |  +-- Lamellidens_corrianus **        
  |  |                          |  |        +--+                                
  |  |                          |  |           +-- Lamellidens_nongyangensis **    
  |  |                          +--+                                            
  |  |                             |           +-- Lamellidens_ferrugineus **      
  |  |                             |  +--------+                                
  |  |                             |  |        +-- Lamellidens_marginalis **       
  |  |                             +--+                                         
  |  |                                |  +-------- Lamellidens_lamellatus **       
  |  |                                +--+                                      
  |  |                                   |  +----- Lamellidens_jenkinsianus **     
  +--+                                   +--+                                   
     |                                      |  +-- Lamellidens_generosus **        
     |                                      +--+                                
     |                                         +-- Lamellidens_mainwaringi **      
     |                                                                          
     |                                +----------- Parreysia_keralaensis **        
     |  +-----------------------------+                                         
     |  |                             |  +-------- Parreysia_corrugata **          
     |  |                             +--+                                      
     |  |                                |  +----- Parreysia_rakhinensis **        
     |  |                                +--+                                   
     |  |                                   |  +-- Parreysia_plagiosoma **         
     |  |                                   +--+                                
     |  |                                      +-- Parreysia_nagpoorensis **       
     |  |                                                                       
     |  |                                      +-- Leoparreysia_burmana **         
     +--+                                +-----+                                
        |                                |     +-- Leoparreysia_whitteni **        
        |  +-----------------------------+                                      
        |  |                             |  +----- Leoparreysia_olivacea **        
        |  |                             +--+                                   
        |  |                                |  +-- Leoparreysia_canefrii **        
        |  |                                +--+                                
        |  |                                   +-- Leoparreysia_tavoyensis **      
        |  |                                                                    
        |  |                             +-------- Nitia_teretiuscula **           
        +--+  +--------------------------+                                      
           |  |                          |  +----- Coelatura_aegyptiaca **         
           |  |                          +--+                                   
           |  |                             |  +-- Coelatura_kunenensis **         
           |  |                             +--+                                
           |  |                                +-- Nyassunio_nyassaensis **        
           |  |                                                                 
           |  |                 +----------------- Indonaia_cylindrica **          
           |  |                 |                                               
           +--+  +--------------+  +-------------- Indonaia_gratiosa **            
              |  |              |  |                                            
              |  |              +--+  +----------- Indonaia_andersoniana **        
              |  |                 |  |                                         
              |  |                 +--+        +-- Indonaia_caerulea **            
              |  |                    |  +-----+                                
              |  |                    |  |     +-- Indonaia_subclathrata **        
              |  |                    +--+                                      
              |  |                       |  +----- Indonaia_rugosa **              
              +--+                       +--+                                   
                 |                          |  +-- Indonaia_bonneaudii **          
                 |                          +--+                                
                 |                             +-- Indonaia_shurtleffiana **       
                 |                                                              
                 |  +----------------------------- Indochinella_pugio **           
                 |  |                                                           
                 |  |                    +-------- Radiatula_crispisulcata **      
                 |  |                    |                                      
                 |  |  +-----------------+     +-- Radiatula_chaudhurii **         
                 +--+  |                 |  +--+                                
                    |  |                 |  |  +-- Radiatula_myitkyinae **         
                    |  |                 +--+                                   
                    |  |                    |  +-- Radiatula_myitthanensis **      
                    |  |                    |  |                                
                    +--+                    +--+-- Radiatula_mouhoti **            
                       |                       |                                
                       |                       +-- Radiatula_chindwinensis **      
                       |                                                        
                       |  +----------------------- Scabiellus_songkramensis **     
                       |  |                                                     
                       +--+  +-------------------- Harmandia_somboriensis **       
                          |  |                                                  
                          +--+  +----------------- Unionetta_fabagina **           
                             |  |                                               
                             +--+              +-- Scabies_crispata **             
                                |  +-----------+                                
                                |  |           +-- Scabies_mandarinus **           
                                +--+                                            
                                   |  +----------- Scabies_nucleus **              
                                   |  |                                         
                                   +--+        +-- Scabies_humilis **              
                                      |  +-----+                                
                                      |  |     +-- Scabies_pilata **               
                                      +--+                                      
                                         |  +----- Scabies_phaselus **             
                                         +--+                                   
                                            |  +-- Scabies_anceps **               
                                            +--+                                
                                               +-- Scabies_scobinatus **           

Bolotov et al., 2022 [Publ 4239], fig. S2

iT tT bi riS oT roT roS iD
58 58 54 2 0 0 0 100.0
                                                  +-- Pseudodon_lenyanensis **        
                                                  |                                
                                                  +-- Nyeinchanconcha_nyeinchani **   
  +-----------------------------------------------+                                
  |                                               +-- Indopseudodon_kayinensis **     
  |                                               |                                
  |                                               +-- Sundadontina_tanintharyiensis **
  |                                                                                
  |                                      +----------- Parreysia_keralaensis **        
  |  +-----------------------------------+                                         
  |  |                                   |  +-------- Parreysia_corrugata **          
  |  |                                   +--+                                      
  |  |                                      |  +----- Parreysia_nagpoorensis **       
  |  |                                      +--+                                   
--+  |                                         |  +-- Parreysia_plagiosoma **         
  |  |                                         +--+                                
  |  |                                            +-- Parreysia_rakhinensis **        
  |  |                                                                             
  |  |                                   +----------- Trapezidens_scutum **           
  |  |                             +-----+                                         
  |  |                             |     |  +-------- Trapezidens_exolescens **       
  |  |                             |     +--+                                      
  |  |                             |        |  +----- Trapezidens_yeti **             
  |  |                             |        +--+                                   
  |  |                             |           |  +-- Trapezidens_dolichorhynchus **  
  +--+                             |           +--+                                
     |  +--------------------------+              +-- Trapezidens_angustior **        
     |  |                          |                                               
     |  |                          |           +----- Lamellidens_savadiensis **      
     |  |                          |  +--------+                                   
     |  |                          |  |        |  +-- Lamellidens_corrianus **        
     |  |                          |  |        +--+                                
     |  |                          |  |           +-- Lamellidens_nongyangensis **    
     |  |                          +--+                                            
     |  |                             |           +-- Lamellidens_marginalis **       
     |  |                             |  +--------+                                
     |  |                             |  |        +-- Lamellidens_ferrugineus **      
     |  |                             +--+                                         
     |  |                                |  +-------- Lamellidens_lamellatus **       
     |  |                                +--+                                      
     |  |                                   |  +----- Lamellidens_jenkinsianus **     
     +--+                                   +--+                                   
        |                                      |  +-- Lamellidens_mainwaringi **      
        |                                      +--+                                
        |                                         +-- Lamellidens_generosus **        
        |                                                                          
        |                                +----------- Leoparreysia_burmana **         
        |  +-----------------------------+                                         
        |  |                             |  +-------- Leoparreysia_whitteni **        
        |  |                             +--+                                      
        |  |                                |  +----- Leoparreysia_olivacea **        
        |  |                                +--+                                   
        |  |                                   |  +-- Leoparreysia_canefrii **        
        |  |                                   +--+                                
        |  |                                      +-- Leoparreysia_tavoyensis **      
        |  |                                                                       
        +--+                                +-------- Nitia_teretiuscula **           
           |  +-----------------------------+                                      
           |  |                             |  +----- Coelatura_aegyptiaca **         
           |  |                             +--+                                   
           |  |                                |  +-- Coelatura_kunenensis **         
           |  |                                +--+                                
           |  |                                   +-- Nyassunio_nyassaensis **        
           |  |                                                                    
           |  |                                   +-- Indonaia_cylindrica **          
           |  |                          +--------+                                
           +--+                          |        +-- Indonaia_gratiosa **            
              |                          |                                         
              |  +-----------------------+     +----- Indonaia_shurtleffiana **       
              |  |                       |  +--+                                   
              |  |                       |  |  |  +-- Indonaia_bonneaudii **          
              |  |                       |  |  +--+                                
              |  |                       +--+     +-- Indonaia_rugosa **              
              |  |                          |                                      
              |  |                          |  +----- Indonaia_andersoniana **        
              |  |                          +--+                                   
              +--+                             |  +-- Indonaia_caerulea **            
                 |                             +--+                                
                 |                                +-- Indonaia_subclathrata **        
                 |                                                                 
                 |  +-------------------------------- Indochinella_pugio **           
                 |  |                                                              
                 |  |                    +----------- Radiatula_crispisulcata **      
                 |  |                    |                                         
                 |  |  +-----------------+        +-- Radiatula_chaudhurii **         
                 +--+  |                 |  +-----+                                
                    |  |                 |  |     +-- Radiatula_myitkyinae **         
                    |  |                 +--+                                      
                    |  |                    |  +----- Radiatula_mouhoti **            
                    |  |                    +--+                                   
                    +--+                       |  +-- Radiatula_myitthanensis **      
                       |                       +--+                                
                       |                          +-- Radiatula_chindwinensis **      
                       |                                                           
                       |  +-------------------------- Scabiellus_songkramensis **     
                       |  |                                                        
                       +--+  +----------------------- Harmandia_somboriensis **       
                          |  |                                                     
                          +--+  +-------------------- Unionetta_fabagina **           
                             |  |                                                  
                             +--+  +----------------- Scabies_crispata **             
                                |  |                                               
                                +--+              +-- Scabies_humilis **              
                                   |  +-----------+                                
                                   |  |           +-- Scabies_mandarinus **           
                                   +--+                                            
                                      |  +----------- Scabies_nucleus **              
                                      +--+                                         
                                         |  +-------- Scabies_pilata **               
                                         +--+                                      
                                            |  +----- Scabies_anceps **               
                                            +--+                                   
                                               |  +-- Scabies_phaselus **             
                                               +--+                                
                                                  +-- Scabies_scobinatus **           

Zanatta & Murphy, 2006 [Publ 2672], fig. 2

iT tT bi riS oT roT roS iD
56 57 48 2 1 1 1 86.0
  +----------------------------------------------------- Pyganodon_grandis **          
  |                                                                                 
  +----------------------------------------------------- Cumberlandia_monodonta     
  |                                                                                 
  |                                            +-------- Pleuronaia_dolabelloides **   
  |                                            |                                    
--+  +-----------------------------------------+-------- Pleurobema_clava **           
  |  |                                         |                                    
  |  |                                         |  +----- Fusconaia_flava **            
  |  |                                         +--+                                 
  |  |                                            |  +-- Hemistena_lata **             
  |  |                                            +--+                              
  |  |                                               +-- Eurynia_dilatata **           
  +--+                                                                              
     |  +----------------------------------------------- Amblema_plicata **            
     |  |                                                                           
     |  +----------------------------------------------- Popenaias_popeii **           
     |  |                                                                           
     |  |                                         +----- Paetulunio_fabalis **         
     |  |                                         |                                 
     |  |  +--------------------------------------+  +-- Toxolasma_parvum **           
     +--+  |                                      |  |                              
        |  |                                      +--+-- Toxolasma_lividum **          
        |  |                                         |                              
        |  |                                         +-- Toxolasma_texasiense **       
        |  |                                                                        
        |  |                                      +----- Obliquaria_reflexa **         
        |  |  +-----------------------------------+                                 
        +--+  |                                   |  +-- Glebula_rotundata **          
           |  |                                   +--+                              
           |  |                                      +-- Cyrtonaias_tampicoensis **    
           |  |                                                                     
           |  |                                   +----- Ptychobranchus_fasciolaris ** 
           |  |                                +--+                                 
           |  |                                |  |  +-- Dromus_dromas **              
           |  |                                |  +--+                              
           +--+  +-----------------------------+     +-- Cyprogenia_stegaria **        
              |  |                             |                                    
              |  |                             |  +----- Medionidus_conradicus **      
              |  |                             +--+                                 
              |  |                                |  +-- Medionidus_acutissimus **     
              |  |                                +--+                              
              |  |                                   +-- Lemiox_rimosus **             
              |  |                                                                  
              |  |  +----------------------------------- Sagittunio_nasutus **         
              |  |  |                                                               
              +--+  |                    +-------------- Potamilus_leptodon **         
                 |  |                    |                                          
                 |  +--------------------+  +----------- Potamilus_fragilis **         
                 |  |                    |  |                                       
                 |  |                    +--+        +-- Potamilus_alatus **           
                 |  |                       |  +-----+                              
                 |  |                       |  |     +-- Potamilus_purpuratus **       
                 |  |                       +--+                                    
                 |  |                          |  +----- Potamilus_inflatus **         
                 |  |                          +--+                                 
                 +--+                             |  +-- Potamilus_ohiensis **         
                    |                             +--+                              
                    |                                +-- Potamilus_amphichaenus **     
                    |                                                               
                    |                                +-- Truncilla_truncata **         
                    |  +-----------------------------+                              
                    |  |                             +-- Ellipsaria_lineolata **       
                    |  |                                                            
                    |  |                             +-- Ortmanniana_pectorosa **      
                    |  |                       +-----+                              
                    |  |                       |     +-- Lampsilis_fasciola **         
                    |  |  +--------------------+                                    
                    +--+  |                    |  +----- Lampsilis_ornata **           
                       |  |                    +--+                                 
                       |  |                       |  +-- Lampsilis_ovata **            
                       |  |                       +--+                              
                       |  |                          +-- Lampsilis_cardium **          
                       |  |                                                         
                       |  |                          +-- Ligumia_recta **              
                       +--+                 +--------+                              
                          |                 |        +-- Cambarunio_iris **            
                          |  +--------------+                                       
                          |  |              |  +-------- Lampsilis_teres **            
                          |  |              |  |                                    
                          |  |              +--+     +-- Hamiota_australis **          
                          |  |                 |  +--+                              
                          |  |                 |  |  +-- Hamiota_subangulata **        
                          +--+                 +--+                                 
                             |                    |  +-- Hamiota_altilis **            
                             |                    +--+                              
                             |                       +-- Hamiota_perovalis **          
                             |                                                      
                             |  +----------------------- Leaunio_vanuxemensis **       
                             |  |                                                   
                             +--+  +-------------------- Ortmanniana_ligamentina **    
                                |  |                                                
                                +--+  +----------------- Lampsilis_siliquoidea **      
                                   |  |                                             
                                   |  |  +-------------- Obovaria_olivaria **          
                                   +--+  |                                          
                                      |  +-------------- Venustaconcha_ellipsiformis **
                                      |  |                                          
                                      |  |        +----- Obovaria_unicolor **          
                                      +--+--------+                                 
                                         |        |  +-- Obovaria_subrotunda **        
                                         |        +--+                              
                                         |           +-- Obovaria_arkansasensis **     
                                         |                                          
                                         |  +----------- Epioblasma_brevidens **       
                                         +--+                                       
                                            |  +-------- Epioblasma_triquetra **       
                                            +--+                                    
                                               |  +----- Epioblasma_rangiana **        
                                               +--+                                 
                                                  |  +-- Epioblasma_walkeri **         
                                                  +--+                              
                                                     +-- Epioblasma_capsaeformis **    

Zanatta & Murphy, 2006 [Publ 2672], fig. 1

iT tT bi riS oT roT roS iD
55 55 50 2 1 1 1 84.0
  +----------------------------------------------------- Cumberlandia_monodonta     
  |                                                                                 
  |  +-------------------------------------------------- Pyganodon_grandis **          
  |  |                                                                              
--+  |                                               +-- Amblema_plicata **            
  |  |                                            +--+                              
  |  |                                            |  +-- Popenaias_popeii **           
  |  |                                            |                                 
  +--+  +-----------------------------------------+  +-- Pleurobema_clava **           
     |  |                                         |  |                              
     |  |                                         |  +-- Eurynia_dilatata **           
     |  |                                         +--+                              
     |  |                                            +-- Pleuronaia_dolabelloides **   
     |  |                                            |                              
     +--+                                            +-- Fusconaia_flava **            
        |                                                                           
        |                                            +-- Cyrtonaias_tampicoensis **    
        |  +-----------------------------------------+                              
        |  |                                         +-- Glebula_rotundata **          
        |  |                                                                        
        |  |  +----------------------------------------- Paetulunio_fabalis **         
        +--+  |                                                                     
           |  |                                +-------- Obliquaria_reflexa **         
           |  |  +-----------------------------+                                    
           |  |  |                             |  +----- Toxolasma_texasiense **       
           |  |  |                             +--+                                 
           +--+  |                                |  +-- Toxolasma_parvum **           
              |  |                                +--+                              
              |  |                                   +-- Toxolasma_lividum **          
              |  |                                                                  
              |  |  +----------------------------------- Sagittunio_nasutus **         
              |  |  |                                                               
              +--+  |                                +-- Medionidus_conradicus **      
                 |  |                          +-----+                              
                 |  |                          |     +-- Ptychobranchus_fasciolaris ** 
                 |  |                 +--------+                                    
                 |  |                 |        |  +----- Medionidus_acutissimus **     
                 |  |                 |        |  |                                 
                 |  |                 |        +--+----- Lemiox_rimosus **             
                 |  |                 |           |                                 
                 |  |  +--------------+           |  +-- Dromus_dromas **              
                 +--+  |              |           +--+                              
                    |  |              |              +-- Cyprogenia_stegaria **        
                    |  |              |                                             
                    |  |              |  +-------------- Potamilus_leptodon **         
                    |  |              |  |                                          
                    |  |              +--+  +----------- Potamilus_fragilis **         
                    |  |                 |  |                                       
                    |  |                 +--+        +-- Potamilus_alatus **           
                    |  |                    |  +-----+                              
                    |  |                    |  |     +-- Potamilus_purpuratus **       
                    |  |                    +--+                                    
                    +--+                       |  +----- Potamilus_inflatus **         
                       |                       +--+                                 
                       |                          |  +-- Potamilus_ohiensis **         
                       |                          +--+                              
                       |                             +-- Potamilus_amphichaenus **     
                       |                                                            
                       |                          +----- Lampsilis_teres **            
                       |  +-----------------------+                                 
                       |  |                       |  +-- Truncilla_truncata **         
                       |  |                       +--+                              
                       |  |                          +-- Ellipsaria_lineolata **       
                       |  |                                                         
                       |  |  +-------------------------- Leaunio_vanuxemensis **       
                       +--+  |                                                      
                          |  |                       +-- Hamiota_australis **          
                          |  |                    +--+                              
                          |  |                    |  +-- Hamiota_subangulata **        
                          |  |              +-----+                                 
                          |  |              |     |  +-- Hamiota_altilis **            
                          |  |              |     +--+                              
                          +--+              |        +-- Hamiota_perovalis **          
                             |  +-----------+                                       
                             |  |           |        +-- Lampsilis_fasciola **         
                             |  |           |  +-----+                              
                             |  |           |  |     +-- Ortmanniana_pectorosa **      
                             |  |           +--+                                    
                             |  |              |  +----- Lampsilis_ornata **           
                             |  |              +--+                                 
                             |  |                 |  +-- Lampsilis_ovata **            
                             +--+                 +--+                              
                                |                    +-- Lampsilis_cardium **          
                                |                                                   
                                |              +-------- Lampsilis_siliquoidea **      
                                |  +-----------+                                    
                                |  |           |  +----- Ortmanniana_ligamentina **    
                                |  |           +--+                                 
                                |  |              |  +-- Cambarunio_iris **            
                                |  |              +--+                              
                                +--+                 +-- Ligumia_recta **              
                                   |                                                
                                   |  +----------------- Venustaconcha_ellipsiformis **
                                   |  |                                             
                                   |  |        +-------- Obovaria_olivaria **          
                                   +--+  +-----+                                    
                                      |  |     |  +----- Obovaria_unicolor **          
                                      |  |     +--+                                 
                                      |  |        |  +-- Obovaria_subrotunda **        
                                      +--+        +--+                              
                                         |           +-- Obovaria_arkansasensis **     
                                         |                                          
                                         |  +----------- Epioblasma_brevidens **       
                                         +--+                                       
                                            |  +-------- Epioblasma_triquetra **       
                                            +--+                                    
                                               |  +----- Epioblasma_rangiana **        
                                               +--+                                 
                                                  |  +-- Epioblasma_walkeri **         
                                                  +--+                              
                                                     +-- Epioblasma_capsaeformis **    

Konopleva et al., 2023 [Publ 4269], fig. S1, 4

iT tT bi riS oT roT roS iD
55 55 53 4 1 1 1 91.0
  +----------------------------------- Gibbosula_laosensis        
  |                                                               
  |                                +-- Trapezidens_exolescens **     
  |                    +-----------+                              
  |                    |           +-- Lamellidens_savadiensis **    
  |  +-----------------+                                          
  |  |                 |  +----------- Parreysia_rakhinensis **      
  |  |                 +--+                                       
  |  |                    |  +-------- Leoparreysia_tavoyensis **    
  |  |                    +--+                                    
  |  |                       |  +----- Indonaia_andersoniana **      
  |  |                       +--+                                 
--+  |                          |  +-- Indochinella_pugio **         
  |  |                          +--+                              
  |  |                             +-- Radiatula_mouhoti **          
  |  |                                                            
  |  |                             +-- Lampsilis_ovata **            
  |  |              +--------------+                              
  |  |              |              +-- Amblema_plicata **            
  |  |              |                                             
  |  |              |     +----------- Sinosolenaia_recognita **     
  |  |              |     |                                       
  |  |              |  +--+        +-- Ptychorhynchus_pfisteri **    
  |  |  +-----------+  |  |  +-----+                              
  +--+  |           |  |  |  |     +-- Parvasolenaia_rivularis **    
     |  |           |  |  +--+                                    
     |  |           |  |     |  +----- Gonidea_angulata **           
     |  |           |  |     +--+                                 
     |  |           |  |        |  +-- Microcondylaea_bonellii **    
     |  |           +--+        +--+                              
     |  |              |           +-- Leguminaia_wheatleyi **       
     |  |              |                                          
     |  |              |        +----- Chamberlainia_somsakpanhai ** 
     |  |              |  +-----+                                 
     |  |              |  |     |  +-- Sinohyriopsis_schlegelii **   
     |  |              |  |     +--+                              
     |  |              +--+        +-- Sinohyriopsis_cumingii **     
     |  |                 |                                       
     |  |                 |  +-------- Potomida_littoralis **        
     |  |                 +--+                                    
     |  |                    |  +----- Pronodularia_japanensis **    
     |  |                    +--+                                 
     |  |                       |  +-- Lamprotula_caveata **         
     +--+                       +--+                              
        |                          +-- Lamprotula_leaii **           
        |                                                         
        |           +----------------- Middendorffinaia_mongolica ** 
        |           |                                             
        |           |        +-------- Aculamprotula_tortuosa **     
        |           |  +-----+                                    
        |  +--------+  |     |  +----- Aculamprotula_scripta **      
        |  |        |  |     +--+                                 
        |  |        |  |        |  +-- Aculamprotula_tientsinensis **
        |  |        |  |        +--+                              
        |  |        +--+           +-- Aculamprotula_coreana **      
        |  |           |                                          
        |  |           |        +----- Unio_tumidus **               
        |  |           |  +-----+                                 
        |  |           |  |     |  +-- Unio_pictorum **              
        |  |           |  |     +--+                              
        |  |           +--+        +-- Unio_crassus **               
        |  |              |                                       
        |  |              |  +-------- Nodularia_douglasiae **       
        +--+              +--+                                    
           |                 |  +----- Tchangsinaia_piscicula **     
           |                 +--+                                 
           |                    |  +-- Cuneopsis_rufescens **        
           |                    +--+                              
           |                       +-- Cuneopsis_heudei **           
           |                                                      
           |                 +-------- Lepidodesma_languilati **     
           |  +--------------+                                    
           |  |              |  +----- Lanceolaria_gladiola **       
           |  |              +--+                                 
           |  |                 |  +-- Lanceolaria_grayii **         
           |  |                 +--+                              
           +--+                    +-- Lanceolaria_triformis **      
              |                                                   
              |  +-------------------- Acuticosta_chinensis **       
              |  |                                                
              |  |              +----- Anodonta_anatina **           
              |  |  +-----------+                                 
              +--+  |           |  +-- Pseudanodonta_complanata **   
                 |  |           +--+                              
                 |  |              +-- Anodonta_cygnea **            
                 |  |                                             
                 |  |        +-------- Anemina_arcaeformis **        
                 +--+  +-----+                                    
                    |  |     |  +----- Amuranodonta_kijaensis **     
                    |  |     +--+                                 
                    |  |        |  +-- Buldowskia_suifunica **       
                    |  |        +--+                              
                    |  |           +-- Buldowskia_shadini **         
                    +--+                                          
                       |     +-------- Beringiana_beringiana **      
                       |  +--+                                    
                       |  |  |  +----- Sinanodonta_schrenkii **      
                       |  |  +--+                                 
                       |  |     |  +-- Sinanodonta_woodiana **       
                       +--+     +--+                              
                          |        +-- Sinanodonta_lauta **          
                          |                                       
                          |  +-------- Cristaria_radiata **          
                          |  |                                    
                          +--+     +-- Cristaria_truncata **         
                             |  +--+                              
                             |  |  +-- Cristaria_plicata **          
                             +--+                                 
                                |  +-- Cristaria_bellua **           
                                +--+                              
                                   +-- Cristaria_clessini **         

Campbell & Lydeard, 2012 [Publ 3308], fig. 3

iT tT bi riS oT roT roS iD
51 51 42 1 0 0 0 79.0
                                   +-------------- Uniomerus_declivis **      
                                   |                                       
  +--------------------------------+           +-- Pustulosa_infucata **      
  |                                |  +--------+                           
  |                                |  |        +-- Pustulosa_kleiniana **     
  |                                +--+                                    
  |                                   |  +-------- Quadrula_quadrula **       
  |                                   +--+                                 
  |                                      |  +----- Cyclonaias_tuberculata **  
  |                                      +--+                              
  |                                         |  +-- Pustulosa_succissa **      
  |                                         +--+                           
  |                                            +-- Pustulosa_pustulosa **     
--+                                                                        
  |                                +-------------- Elliptoideus_sloatianus ** 
  |  +-----------------------------+                                       
  |  |                             |  +----------- Plectomerus_dombeyanus **  
  |  |                             +--+                                    
  |  |                                |  +-------- Obovaria_unicolor **       
  |  |                                +--+                                 
  |  |                                   |  +----- Obovaria_olivaria **       
  |  |                                   +--+                              
  |  |                                      |  +-- Lampsilis_ovata **         
  |  |                                      +--+                           
  |  |                                         +-- Lemiox_rimosus **          
  +--+                                                                     
     |                                +----------- Cyrtonaias_tampicoensis ** 
     |                                |                                    
     |                                |     +----- Popenaias_popeii **        
     |  +-----------------------------+  +--+                              
     |  |                             |  |  |  +-- Amblema_elliottii **       
     |  |                             |  |  +--+                           
     |  |                             +--+     +-- Amblema_plicata **         
     |  |                                |                                 
     |  |                                |     +-- Reginaia_ebenus[1] **      
     |  |                                |     |                           
     +--+                                +-----+-- Reginaia_ebenus[2] **      
        |                                      |                           
        |                                      +-- Reginaia_rotulata **       
        |                                                                  
        |                                      +-- Elliptio_arca **           
        |  +-----------------------------------+                           
        |  |                                   +-- Elliptio_crassidens **     
        |  |                                                               
        +--+  +----------------------------------- Eurynia_dilatata **        
           |  |                                                            
           |  |  +-------------------------------- Parvaspina_collina **      
           |  |  |                                                         
           +--+  |                             +-- Hemistena_lata **          
              |  +-----------------------------+                           
              |  |                             +-- Plethobasus_cyphyus **     
              |  |                                                         
              +--+                          +----- Pleuronaia_barnesiana **   
                 |  +-----------------------+                              
                 |  |                       |  +-- Pleuronaia_dolabelloides **
                 |  |                       +--+                           
                 |  |                          +-- Pleuronaia_gibber **       
                 |  |                                                      
                 |  |                          +-- Pleurobema_stabile **      
                 +--+                    +-----+                           
                    |                    |     +-- Pleurobema_sintoxia **     
                    |  +-----------------+                                 
                    |  |                 |  +----- Pleurobema_decisum **      
                    |  |                 +--+                              
                    |  |                    |  +-- Pleurobema_clava **        
                    +--+                    +--+                           
                       |                       +-- Pleurobema_pyriforme **    
                       |                                                   
                       |  +----------------------- Pleurobema_cordatum **     
                       |  |                                                
                       +--+  +-------------------- Fusconaia_subrotunda **    
                          |  |                                             
                          |  |                 +-- Fusconaia_cor **           
                          +--+  +--------------+                           
                             |  |              +-- Fusconaia_cuneolus **      
                             |  |                                          
                             +--+  +-------------- Fusconaia_ozarkensis **    
                                |  |                                       
                                |  |           +-- Fusconaia_flava[6] **      
                                |  |  +--------+                           
                                +--+  |        +-- Fusconaia_masoni **        
                                   |  |                                    
                                   |  |        +-- Fusconaia_escambia **      
                                   |  |        |                           
                                   +--+        +-- Fusconaia_burkei[1] **     
                                      |  +-----+                           
                                      |  |     +-- Fusconaia_burkei[2] **     
                                      |  |     |                           
                                      |  |     +-- Fusconaia_burkei[3] **     
                                      +--+                                 
                                         |  +----- Fusconaia_flava[5] **      
                                         |  |                              
                                         |  |  +-- Fusconaia_cerina **        
                                         +--+  |                           
                                            |  +-- Fusconaia_flava[1] **      
                                            |  |                           
                                            +--+-- Fusconaia_flava[2] **      
                                               |                           
                                               +-- Fusconaia_flava[3] **      
                                               |                           
                                               +-- Fusconaia_flava[4] **      

Wu et al., 2018 [Publ 3836], fig. 2

iT tT bi riS oT roT roS iD
51 51 43 3 2 2 1 85.0
                                   +-- Margaritifera_dahurica     
  +--------------------------------+                              
  |                                +-- Margaritifera_falcata      
  |                                                               
  |  +-------------------------------- Quadrula_quadrula **          
  |  |                                                            
  |  |                             +-- Amblema_plicata **            
  |  +-----------------------------+                              
  |  |                             +-- Obovaria_arkansasensis **     
  |  |                                                            
--+  |                    +----------- Pleurobema_sintoxia **        
  |  |                    |                                       
  |  +--------------------+        +-- Elliptio_complanata **        
  |  |                    |  +-----+                              
  |  |                    |  |     +-- Eurynia_dilatata **           
  |  |                    +--+                                    
  |  |                       |  +----- Fusconaia_askewi **           
  |  |                       +--+                                 
  |  |                          |  +-- Fusconaia_cerina **           
  |  |                          +--+                              
  +--+                             +-- Fusconaia_burkei **           
     |                                                            
     |                          +----- Potomida_littoralis **        
     |                    +-----+                                 
     |                    |     |  +-- Lamprotula_caveata **         
     |                    |     +--+                              
     |  +-----------------+        +-- Lamprotula_leaii **           
     |  |                 |                                       
     |  |                 |  +-------- Sinosolenaia_carinata **      
     |  |                 |  |                                    
     |  |                 +--+  +----- Gonidea_angulata **           
     |  |                    |  |                                 
     |  |                    +--+----- Sinosolenaia_oleivora **      
     |  |                       |                                 
     |  |                       |  +-- Parvasolenaia_rivularis **    
     |  |                       +--+                              
     |  |                          +-- Ptychorhynchus_pfisteri **    
     +--+                                                         
        |  +-------------------------- Sinohyriopsis_cumingii **     
        |  |                                                      
        |  |  +----------------------- Lepidodesma_languilati **     
        |  |  |                                                   
        |  |  |           +----------- Aculamprotula_tortuosa **     
        |  |  |     +-----+                                       
        |  |  |     |     |  +-------- Aculamprotula_scripta **      
        |  |  |     |     +--+                                    
        |  |  |     |        |  +----- Aculamprotula_coreana **      
        |  |  |     |        +--+                                 
        +--+  |     |           |  +-- Aculamprotula_fibrosa **      
           |  +-----+           +--+                              
           |  |     |              +-- Aculamprotula_tientsinensis **
           |  |     |                                             
           |  |     |              +-- Unio_crassus **               
           |  |     |  +-----------+                              
           |  |     |  |           +-- Unio_pictorum **              
           |  |     |  |                                          
           |  |     +--+        +----- Nodularia_douglasiae **       
           |  |        |  +-----+                                 
           |  |        |  |     |  +-- Schistodesmus_lampreyanus **  
           +--+        |  |     +--+                              
              |        +--+        +-- Schistodesmus_spinosus **     
              |           |                                       
              |           |  +-------- Tchangsinaia_piscicula **     
              |           +--+                                    
              |              |  +----- Cuneopsis_rufescens **        
              |              +--+                                 
              |                 |  +-- Cuneopsis_celtiformis **      
              |                 +--+                              
              |                    +-- Cuneopsis_heudei **           
              |                                                   
              |           +----------- Acuticosta_chinensis **       
              |  +--------+                                       
              |  |        |  +-------- Lanceolaria_gladiola **       
              |  |        +--+                                    
              |  |           |  +----- Lanceolaria_grayii **         
              |  |           +--+                                 
              |  |              |  +-- Lanceolaria_lanceolata **     
              |  |              +--+                              
              |  |                 +-- Lanceolaria_triformis **      
              +--+                                                
                 |           +-------- Anemina_arcaeformis **        
                 |           |                                    
                 |           +-------- Cristaria_plicata **          
                 |  +--------+                                    
                 |  |        |     +-- Sinanodonta_woodiana **       
                 |  |        |  +--+                              
                 |  |        |  |  +-- Sinanodonta_elliptica **      
                 |  |        +--+                                 
                 +--+           |  +-- Sinanodonta_lucida **         
                    |           +--+                              
                    |              +-- Sinanodonta_angula **         
                    |                                             
                    |  +-------------- Anodonta_cygnea **            
                    |  |                                          
                    +--+  +----------- Anodonta_anatina **           
                       |  |                                       
                       +--+  +-------- Strophitus_undulatus **       
                          |  |                                    
                          +--+  +----- Platynaias_compressa **       
                             |  |                                 
                             +--+----- Alasmidonta_marginata **      
                                |                                 
                                |  +-- Pyganodon_cataracta **        
                                +--+                              
                                   +-- Pyganodon_grandis **          

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iT tT bi riS oT roT roS iD
51 51 49 3 3 3 1 86.0
                          +----- Gibbosula_crassa           
  +-----------------------+                                 
  |                       |  +-- Pseudunio_marocanus        
  |                       +--+                              
  |                          +-- Cumberlandia_monodonta     
  |                                                         
  |                          +-- Quadrula_quadrula **          
  |        +-----------------+                              
  |        |                 +-- Uniomerus_tetralasmus **      
  |     +--+                                                
  |     |  |  +----------------- Elliptio_complanata **        
  |     |  |  |                                             
  |     |  +--+              +-- Potamilus_leptodon **         
  |     |     |  +-----------+                              
  |     |     |  |           +-- Potamilus_alatus **           
  |     |     +--+                                          
  |     |        |  +----------- Leaunio_lienosus **           
  |     |        |  |                                       
  |     |        +--+        +-- Lampsilis_cardium **          
  |     |           |  +-----+                              
--+     |           |  |     +-- Lampsilis_ornata **           
  |  +--+           +--+                                    
  |  |  |              |  +----- Venustaconcha_ellipsiformis **
  |  |  |              +--+                                 
  |  |  |                 |  +-- Lampsilis_powellii **         
  |  |  |                 +--+                              
  |  |  |                    +-- Lampsilis_siliquoidea **      
  |  |  |                                                   
  |  |  |                    +-- Sinohyriopsis_cumingii **     
  |  |  |        +-----------+                              
  |  |  |        |           +-- Sinohyriopsis_schlegelii **   
  |  |  |        |                                          
  |  |  |        |           +-- Pilsbryoconcha_exilis **      
  |  |  +--------+  +--------+                              
  |  |           |  |        +-- Pseudodon_vondembuschianus ** 
  |  |           |  |                                       
  |  |           |  |        +-- Lamprotula_caveata **         
  |  |           |  |     +--+                              
  |  |           +--+     |  +-- Lamprotula_leaii **           
  |  |              |  +--+                                 
  +--+              |  |  |  +-- Pronodularia_japanensis **    
     |              |  |  +--+                              
     |              |  |     +-- Potomida_littoralis **        
     |              +--+                                    
     |                 |     +-- Sinosolenaia_carinata **      
     |                 |  +--+                              
     |                 |  |  +-- Sinosolenaia_oleivora **      
     |                 +--+                                 
     |                    |  +-- Parvasolenaia_rivularis **    
     |                    +--+                              
     |                       +-- Ptychorhynchus_pfisteri **    
     |                                                      
     |  +----------------------- Lepidodesma_languilati **     
     |  |                                                   
     |  |                 +----- Lanceolaria_gladiola **       
     |  |  +--------------+                                 
     |  |  |              |  +-- Lanceolaria_grayii **         
     |  |  |              +--+                              
     |  |  |                 +-- Lanceolaria_lanceolata **     
     +--+  |                                                
        |  |     +-------------- Acuticosta_chinensis **       
        |  |     |                                          
        |  |     |     +-------- Cristaria_plicata **          
        |  |     |     |                                    
        |  |     |  +--+     +-- Sinanodonta_woodiana **       
        |  |  +--+  |  |  +--+                              
        +--+  |  |  |  |  |  +-- Sinanodonta_lucida **         
           |  |  |  |  +--+                                 
           |  |  |  |     |  +-- Anemina_arcaeformis **        
           |  |  |  |     +--+                              
           |  |  +--+        +-- Anemina_euscaphys **          
           |  |     |                                       
           |  |     |        +-- Anodonta_anatina **           
           |  |     |  +-----+                              
           |  |     |  |     +-- Anodonta_cygnea **            
           |  |     |  |                                    
           |  |     +--+     +-- Pyganodon_grandis **          
           +--+        |  +--+                              
              |        |  |  +-- Utterbackia_peninsularis **   
              |        +--+                                 
              |           |  +-- Alasmidonta_varicosa **       
              |           +--+                              
              |              +-- Prolasmidonta_heterodon **    
              |                                             
              |           +----- Aculamprotula_tortuosa **     
              |  +--------+                                 
              |  |        |  +-- Aculamprotula_tientsinensis **
              |  |        +--+                              
              |  |           +-- Aculamprotula_coreana **      
              |  |                                          
              +--+     +-------- Unio_tumidus **               
                 |  +--+                                    
                 |  |  |  +----- Unio_crassus **               
                 |  |  +--+                                 
                 |  |     |  +-- Unio_pictorum **              
                 |  |     +--+                              
                 +--+        +-- Unio_delphinus **             
                    |                                       
                    |        +-- Schistodesmus_lampreyanus **  
                    |  +-----+                              
                    |  |     +-- Cuneopsis_heudei **           
                    +--+                                    
                       |  +----- Pseudocuneopsis_capitata **   
                       +--+                                 
                          |  +-- Nodularia_douglasiae **       
                          +--+                              
                             +-- Tchangsinaia_piscicula **     

Dai et al., 2023 [Publ 4285], fig. 5

iT tT bi riS oT roT roS iD
51 51 49 3 2 2 1 87.0
                                   +-- Gibbosula_rochechouartii   
  +--------------------------------+                              
  |                                +-- Margaritifera_dahurica     
  |                                                               
  |                                +-- Quadrula_quadrula **          
  |     +--------------------------+                              
  |     |                          +-- Uniomerus_tetralasmus **      
  |     |                                                         
  |  +--+                          +-- Elliptio_complanata **        
  |  |  |  +-----------------------+                              
  |  |  |  |                       +-- Pleurobema_oviforme **        
  |  |  +--+                                                      
  |  |     |  +----------------------- Amblema_plicata **            
  |  |     |  |                                                   
  |  |     +--+  +-------------------- Popenaias_popeii **           
  |  |        |  |                                                
--+  |        +--+  +----------------- Toxolasma_parvum **           
  |  |           |  |                                             
  |  |           +--+              +-- Potamilus_leptodon **         
  |  |              |  +-----------+                              
  |  |              |  |           +-- Potamilus_alatus **           
  |  |              +--+                                          
  |  |                 |  +----------- Leaunio_lienosus **           
  |  |                 |  |                                       
  |  |                 +--+        +-- Lampsilis_cardium **          
  |  |                    |  +-----+                              
  |  |                    |  |     +-- Lampsilis_ornata **           
  |  |                    +--+                                    
  |  |                       |  +----- Venustaconcha_ellipsiformis **
  |  |                       +--+                                 
  |  |                          |  +-- Lampsilis_powellii **         
  |  |                          +--+                              
  +--+                             +-- Lampsilis_siliquoidea **      
     |                                                            
     |           +-------------------- Lepidodesma_languilati **     
     |           |                                                
     |           |     +-------------- Aculamprotula_tientsinensis **
     |           |     |                                          
     |           |  +--+  +----------- Unio_crassus **               
     |           |  |  |  |                                       
     |        +--+  |  +--+        +-- Pseudocuneopsis_capitata **   
     |        |  |  |     |  +-----+                              
     |        |  |  |     |  |     +-- Nodularia_douglasiae **       
     |        |  |  |     +--+                                    
     |        |  |  |        |  +----- Schistodesmus_lampreyanus **  
     |        |  |  |        +--+                                 
     |        |  +--+           |  +-- Cuneopsis_celtiformis **      
     |        |     |           +--+                              
     |        |     |              +-- Tchangsinaia_piscicula **     
     |        |     |                                             
     |        |     |              +-- Acuticosta_chinensis **       
     |        |     |  +-----------+                              
     |        |     |  |           +-- Lanceolaria_grayii **         
     |        |     |  |                                          
     |        |     +--+        +----- Cristaria_plicata **          
     |        |        |  +-----+                                 
     |        |        |  |     |  +-- Anemina_arcaeformis **        
     +--------+        |  |     +--+                              
              |        +--+        +-- Sinanodonta_woodiana **       
              |           |                                       
              |           |  +-------- Anodonta_cygnea **            
              |           +--+                                    
              |              |  +----- Platynaias_compressa **       
              |              +--+                                 
              |                 |  +-- Pyganodon_grandis **          
              |                 +--+                              
              |                    +-- Utterbackia_imbecillis **     
              |                                                   
              |                 +----- Chamberlainia_somsakpanhai ** 
              |  +--------------+                                 
              |  |              |  +-- Sinohyriopsis_cumingii **     
              |  |              +--+                              
              |  |                 +-- Sinohyriopsis_schlegelii **   
              |  |                                                
              |  |                 +-- Hyriopsis_bialata **          
              +--+              +--+                              
                 |              |  +-- Rectidens_sumatrensis **      
                 |  +-----------+                                 
                 |  |           |  +-- Lens_contradens **            
                 |  |           +--+                              
                 |  |              +-- Physunio_superbus **          
                 |  |                                             
                 +--+        +-------- Potomida_littoralis **        
                    |  +-----+                                    
                    |  |     |  +----- Pronodularia_japanensis **    
                    |  |     +--+                                 
                    |  |        |  +-- Lamprotula_caveata **         
                    |  |        +--+                              
                    +--+           +-- Lamprotula_leaii **           
                       |                                          
                       |           +-- Pseudodon_vondembuschianus ** 
                       |  +--------+                              
                       |  |        +-- Pilsbryoconcha_exilis **      
                       |  |                                       
                       +--+     +----- Parvasolenaia_rivularis **    
                          |  +--+                                 
                          |  |  |  +-- Postolata_guangxiensis **     
                          |  |  +--+                              
                          +--+     +-- Ptychorhynchus_pfisteri **    
                             |                                    
                             |  +----- Microcondylaea_bonellii **    
                             +--+                                 
                                |  +-- Sinosolenaia_oleivora **      
                                +--+                              
                                   +-- Sinosolenaia_carinata **      

Konopleva et al., 2019 [Publ 3900], fig. 1

iT tT bi riS oT roT roS iD
50 50 39 2 2 2 1 90.0
                                   +-- Gibbosula_laosensis        
  +--------------------------------+                              
  |                                +-- Margaritifera_dahurica     
  |                                                               
  |                                +-- Indopseudodon_indawgyiensis **
  |                                |                              
  |                             +--+-- Indopseudodon_bogani **       
  |                             |  |                              
  |                             |  +-- Indopseudodon_salwenianus **  
  |                             |                                 
  |                             |  +-- Bineurus_mouhotii **          
  |  +--------------------------+  |                              
  |  |                          |  +-- Pseudodon_vagulus **          
  |  |                          |  |                              
  |  |                          |  +-- Pseudodon_mekongi **          
--+  |                          |  |                              
  |  |                          |  +-- Pilsbryoconcha_exilis **      
  |  |                          +--+                              
  |  |                             +-- Pilsbryoconcha_carinifera **  
  |  |                             |                              
  |  |                             +-- Bineurus_loeiensis **         
  |  |                             |                              
  |  |                             +-- Bineurus_exilis **            
  |  |                             |                              
  |  |                             +-- Pseudodon_vondembuschianus ** 
  |  |                                                            
  |  |                    +----------- Indonaia_shurtleffiana **     
  |  |                 +--+                                       
  |  |                 |  |  +-------- Indonaia_rugosa **            
  |  |                 |  +--+                                    
  +--+                 |     |  +----- Indonaia_andersoniana **      
     |                 |     +--+                                 
     |                 |        |  +-- Indonaia_caerulea **          
     |  +--------------+        +--+                              
     |  |              |           +-- Indonaia_subclathrata **      
     |  |              |                                          
     |  |              |  +----------- Indochinella_pugio **         
     |  |              |  |                                       
     |  |              |  |        +-- Scabies_humilis **            
     |  |              +--+        |                              
     |  |                 |  +-----+-- Scabies_mandarinus **         
     |  |                 |  |     |                              
     |  |                 +--+     +-- Scabies_phaselus **           
     |  |                    |                                    
     |  |                    |  +----- Radiatula_mouhoti **          
     |  |                    +--+                                 
     |  |                       |  +-- Radiatula_myitkyinae **       
     |  |                       +--+                              
     |  |                          +-- Radiatula_chaudhurii **       
     |  |                                                         
     +--+                          +-- Ensidens_ingallsianus **      
        |                    +-----+                              
        |                    |     +-- Ensidens_telus **             
        |              +-----+                                    
        |              |     |  +----- Ensidens_sagittarius **       
        |              |     +--+                                 
        |              |        |  +-- Ensidens_jaculus **           
        |  +-----------+        +--+                              
        |  |           |           +-- Ensidens_spiculus **          
        |  |           |                                          
        |  |           |  +----------- Hyriopsis_desowitzi **        
        |  |           |  |                                       
        |  |           +--+----------- Rectidens_sumatrensis **      
        |  |              |                                       
        |  |              |  +-------- Hyriopsis_panhai **           
        |  |              +--+                                    
        |  |                 |  +----- Hyriopsis_myersiana **        
        |  |                 +--+                                 
        +--+                    |  +-- Hyriopsis_khoratensis **      
           |                    +--+                              
           |                       +-- Hyriopsis_bialata **          
           |                                                      
           |                       +-- Yaukthwa_paiensis **          
           |                 +-----+                              
           |                 |     +-- Yaukthwa_inlenensis **        
           |  +--------------+                                    
           |  |              |  +----- Yaukthwa_panhai **            
           |  |              +--+                                 
           |  |                 |  +-- Yaukthwa_nesemanni **         
           |  |                 +--+                              
           +--+                    +-- Yaukthwa_rectangularis **     
              |                                                   
              |                 +----- Physunio_superbus **          
              |  +--------------+                                 
              |  |              |  +-- Physunio_modelli **           
              |  |              +--+                              
              +--+                 +-- Physunio **                   
                 |                                                
                 |  +----------------- Trapezoideus_foliaceus **     
                 |  |                                             
                 +--+              +-- Lens_contradens **            
                    |  +-----------+                              
                    |  |           +-- Lens_micropterus **           
                    +--+                                          
                       |  +----------- Lens_misellus **              
                       +--+                                       
                          |  +-------- Lens_eximius **               
                          +--+                                    
                             |  +----- Lens_pallegoixi **            
                             +--+                                 
                                |  +-- Lens_comptus **               
                                +--+                              
                                   +-- Lens_novoselovi **            

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT tT bi riS oT roT roS iD
50 50 48 4 7 2 1 88.0
                                +-- Margaritifera_margaritifera  
  +-----------------------------+                                
  |                             +-- Margaritifera_hembeli        
  |                                                              
  |                             +-- Quadrula_quadrula **            
  |        +--------------------+                                
  |        |                    +-- Uniomerus_tetralasmus **        
  |        |                                                     
  |        |                 +----- Elliptoideus_sloatianus **      
  |  +-----+  +--------------+                                   
  |  |     |  |              |  +-- Elliptio_complanata **          
  |  |     |  |              +--+                                
  |  |     |  |                 +-- Pleurobema_strodeanum **        
  |  |     |  |                                                  
  |  |     +--+              +----- Reginaia_ebenus **              
  |  |        |  +-----------+                                   
  |  |        |  |           |  +-- Plectomerus_dombeyanus **       
--+  |        |  |           +--+                                
  |  |        |  |              +-- Amblema_plicata **              
  |  |        +--+                                               
  |  |           |              +-- Psoronaias_semigranosa **       
  |  |           |  +-----------+                                
  |  |           |  |           +-- Popenaias_popeii **             
  |  |           +--+                                            
  |  |              |  +----------- Obliquaria_reflexa **           
  |  |              +--+                                         
  |  |                 |  +-------- Cyrtonaias_tampicoensis **      
  |  |                 +--+                                      
  |  |                    |  +----- Leaunio_lienosus **             
  |  |                    +--+                                   
  |  |                       |  +-- Lampsilis_cardium **            
  |  |                       +--+                                
  |  |                          +-- Pachynaias_spheniopsis **       
  |  |                                                           
  +--+                          +-- Chamberlainia_hainesiana **     
     |     +--------------------+                                
     |     |                    +-- Sinohyriopsis_cumingii **       
     |     |                                                     
     |     |     +----------------- Lamprotula_cornuumlunae **      
     |     |  +--+                                               
     |     |  |  |  +-------------- Gonidea_angulata **             
     |  +--+  |  +--+                                            
     |  |  |  |     |  +----------- Indopseudodon_crebristriatus ** 
     |  |  |  |     +--+                                         
     |  |  |  |        |  +-------- Bineurus_anodontinum **         
     |  |  |  |        +--+                                      
     |  |  |  |           |  +----- Pilsbryoconcha_exilis **        
     |  |  |  |           +--+                                   
     |  |  +--+              |  +-- Pseudodon_vondembuschianus **   
     |  |     |              +--+                                
     |  |     |                 +-- Pseudodon_inoscularis **        
     |  |     |                                                  
     |  |     |              +----- Ensidens_ingallsianus **        
     |  |     |        +-----+                                   
     |  |     |        |     |  +-- Hyriopsis_bialata **            
     |  |     |        |     +--+                                
     |  |     |        |        +-- Rectidens_sumatrensis **        
     |  |     +--------+                                         
     |  |              |        +-- Trapezoideus_foliaceus **       
     |  |              |  +-----+                                
     +--+              |  |     +-- Physunio_superbus **            
        |              +--+                                      
        |                 |  +----- Yaukthwa_paiensis **            
        |                 +--+                                   
        |                    |  +-- Lens_contradens **              
        |                    +--+                                
        |                       +-- Solenaia_emarginata **          
        |                                                        
        |              +----------- Lanceolaria_lanceolata **       
        |              |                                         
        |              |     +----- Unio_gibbus **                  
        |     +--------+  +--+                                   
        |     |        |  |  |  +-- Nodularia_douglasiae **         
        |     |        |  |  +--+                                
        |     |        |  |     +-- Tchangsinaia_piscicula **       
        |     |        +--+                                      
        |     |           |     +-- Cristaria_plicata **            
        |     |           |  +--+                                
        |     |           |  |  +-- Sinanodonta_woodiana **         
        +-----+           +--+                                   
              |              |  +-- Anodonta_cygnea **              
              |              +--+                                
              |                 +-- Pyganodon_grandis **            
              |                                                  
              |                 +-- Trapezidens_dolichorhynchus **  
              |  +--------------+                                
              |  |              +-- Lamellidens_savadiensis **      
              |  |                                               
              +--+              +-- Leoparreysia_tavoyensis **      
                 |  +-----------+                                
                 |  |           +-- Leoparreysia_olivacea **        
                 |  |                                            
                 +--+           +-- Coelatura_choziensis **         
                    |  +--------+                                
                    |  |        +-- Prisodontopsis_aviculaeformis **
                    +--+                                         
                       |  +-------- Indonaia_caerulea **            
                       +--+                                      
                          |  +----- Indochinella_pugio **           
                          +--+                                   
                             |  +-- Scabies_phaselus **             
                             +--+                                
                                +-- Harmandia_somboriensis **       

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT tT bi riS oT roT roS iD
50 50 47 4 7 2 1 85.0
                       +-- Quadrula_quadrula **            
     +-----------------+                                
     |                 +-- Uniomerus_tetralasmus **        
     |                                                  
     |              +----- Elliptoideus_sloatianus **      
     |           +--+                                   
     |           |  |  +-- Elliptio_complanata **          
  +--+           |  +--+                                
  |  |  +--------+     +-- Pleurobema_strodeanum **        
  |  |  |        |                                      
  |  |  |        |  +----- Reginaia_ebenus **              
  |  |  |        +--+                                   
  |  |  |           |  +-- Plectomerus_dombeyanus **       
  |  +--+           +--+                                
  |     |              +-- Amblema_plicata **              
  |     |                                               
  |     |              +-- Psoronaias_semigranosa **       
  |     |  +-----------+                                
  |     |  |           +-- Popenaias_popeii **             
  |     +--+                                            
  |        |  +----------- Obliquaria_reflexa **           
  |        +--+                                         
  |           |  +-------- Cyrtonaias_tampicoensis **      
  |           +--+                                      
  |              |  +----- Leaunio_lienosus **             
  |              +--+                                   
  |                 |  +-- Lampsilis_cardium **            
  |                 +--+                                
  |                    +-- Pachynaias_spheniopsis **       
  |                                                     
  |                    +-- Chamberlainia_hainesiana **     
  |  +-----------------+                                
  |  |                 +-- Sinohyriopsis_cumingii **       
  |  |                                                  
  |  |                 +-- Gonidea_angulata **             
  |  |     +-----------+                                
  |  |     |           +-- Lamprotula_cornuumlunae **      
  |  |  +--+                                            
  +--+  |  |  +----------- Indopseudodon_crebristriatus ** 
  |  |  |  +--+                                         
  |  |  |     |  +-------- Pilsbryoconcha_exilis **        
  |  |  |     +--+                                      
--+  |  |        |  +----- Bineurus_anodontinum **         
  |  |  |        +--+                                   
  |  |  |           |  +-- Pseudodon_vondembuschianus **   
  |  +--+           +--+                                
  |     |              +-- Pseudodon_inoscularis **        
  |     |                                               
  |     |           +----- Ensidens_ingallsianus **        
  |     |     +-----+                                   
  |     |     |     |  +-- Hyriopsis_bialata **            
  |     |     |     +--+                                
  |     |     |        +-- Rectidens_sumatrensis **        
  |     +-----+                                         
  |           |        +-- Trapezoideus_foliaceus **       
  |           |  +-----+                                
  |           |  |     +-- Physunio_superbus **            
  |           +--+                                      
  |              |  +----- Yaukthwa_paiensis **            
  |              +--+                                   
  |                 |  +-- Lens_contradens **              
  |                 +--+                                
  |                    +-- Solenaia_emarginata **          
  |                                                     
  |              +-------- Lanceolaria_lanceolata **       
  |           +--+                                      
  |           |  |  +----- Unio_gibbus **                  
  |           |  +--+                                   
  |           |     |  +-- Tchangsinaia_piscicula **       
  |           |     +--+                                
  |  +--------+        +-- Nodularia_douglasiae **         
  |  |        |                                         
  |  |        |        +-- Cristaria_plicata **            
  |  |        |     +--+                                
  |  |        |     |  +-- Sinanodonta_woodiana **         
  |  |        +-----+                                   
  |  |              |  +-- Anodonta_cygnea **              
  +--+              +--+                                
     |                 +-- Pyganodon_grandis **            
     |                                                  
     |                 +-- Trapezidens_dolichorhynchus **  
     |  +--------------+                                
     |  |              +-- Lamellidens_savadiensis **      
     |  |                                               
     +--+              +-- Leoparreysia_tavoyensis **      
        |  +-----------+                                
        |  |           +-- Leoparreysia_olivacea **        
        |  |                                            
        +--+           +-- Coelatura_choziensis **         
           |  +--------+                                
           |  |        +-- Prisodontopsis_aviculaeformis **
           +--+                                         
              |  +-------- Indonaia_caerulea **            
              +--+                                      
                 |  +----- Indochinella_pugio **           
                 +--+                                   
                    |  +-- Scabies_phaselus **             
                    +--+                                
                       +-- Harmandia_somboriensis **       

Sano et al., 2020 [Publ 4138], fig. 2

iT tT bi riS oT roT roS iD
50 50 48 4 23 12 1 83.0
                          +-------------------- Cumberlandia_monodonta     
                          |                                                
                          |                 +-- Gibbosula_rochechouartii   
                          |              +--+                              
  +-----------------------+              |  +-- Gibbosula_laosensis        
  |                       |  +-----------+                                 
  |                       |  |           |  +-- Pseudunio_auricularius     
  |                       |  |           +--+                              
  |                       |  |              +-- Pseudunio_marocanus        
  |                       +--+                                             
  |                          |              +-- Margaritifera_dahurica     
  |                          |  +-----------+                              
  |                          |  |           +-- Margaritifera_margaritifera
  |                          +--+                                          
  |                             |  +----------- Margaritifera_falcata      
  |                             +--+                                       
  |                                |  +-------- Margaritifera_middendorffi 
  |                                +--+                                    
--+                                   |  +----- Margaritifera_laevis       
  |                                   +--+                                 
  |                                      |  +-- Margaritifera_hembeli      
  |                                      +--+                              
  |                                         +-- Margaritifera_marrianae    
  |                                                                        
  |  +----------------------------------------- Scabies_crispata **           
  |  |                                                                     
  |  |                       +----------------- Lampsilis_cardium **          
  |  |                       |                                             
  |  |                       |           +----- Sinohyriopsis_cumingii[2] **  
  |  |  +--------------------+  +--------+                                 
  |  |  |                    |  |        |  +-- Sinohyriopsis_schlegelii **   
  |  |  |                    |  |        +--+                              
  |  |  |                    +--+           +-- Sinohyriopsis_cumingii[1] **  
  +--+  |                       |                                          
     |  |                       |  +----------- Inversidens_brandtii **       
     |  |                       |  |                                       
     |  |                       +--+        +-- Ptychorhynchus_pfisteri **    
     |  |                          |  +-----+                              
     |  |                          |  |     +-- Obovalis_omiensis **          
     |  |                          +--+                                    
     |  |                             |  +----- Lamprotula_leaii **           
     |  |                             +--+                                 
     +--+                                |  +-- Pronodularia_japanensis **    
        |                                +--+                              
        |                                   +-- Potomida_littoralis **        
        |                                                                  
        |                                +----- Buldowskia_iwakawai **        
        |  +-----------------------------+                                 
        |  |                             |  +-- Anemina_arcaeformis **        
        |  |                             +--+                              
        |  |                                +-- Anemina_euscaphys **          
        |  |                                                               
        |  |                    +-------------- Pletholophus_tenuis **        
        |  |                    |                                          
        |  |                    |        +----- Beringiana_fukuharai **       
        |  |  +-----------------+  +-----+                                 
        +--+  |                 |  |     |  +-- Beringiana_japonica **        
           |  |                 |  |     +--+                              
           |  |                 +--+        +-- Beringiana_beringiana **      
           |  |                    |                                       
           |  |                    |  +-------- Sinanodonta_lucida **         
           |  |                    +--+                                    
           |  |                       |  +----- Sinanodonta_tumens **         
           |  |                       +--+                                 
           |  |                          |  +-- Sinanodonta_lauta **          
           +--+                          +--+                              
              |                             +-- Sinanodonta_woodiana **       
              |                                                            
              |  +----------------------------- Cristaria_plicata **          
              |  |                                                         
              |  |                          +-- Anodonta_cygnea **            
              |  |                 +--------+                              
              |  |                 |        +-- Anodonta_anatina **           
              |  |                 |                                       
              |  |  +--------------+     +----- Alasmidonta_varicosa **       
              +--+  |              |  +--+                                 
                 |  |              |  |  |  +-- Prolasmidonta_heterodon **    
                 |  |              |  |  +--+                              
                 |  |              +--+     +-- Platynaias_compressa **       
                 |  |                 |                                    
                 |  |                 |  +----- Pyganodon_grandis **          
                 |  |                 +--+                                 
                 +--+                    |  +-- Utterbackia_imbecillis **     
                    |                    +--+                              
                    |                       +-- Utterbackia_peninsularis **   
                    |                                                      
                    |                    +----- Lanceolaria_grayii[1] **      
                    |  +-----------------+                                 
                    |  |                 |  +-- Lanceolaria_lanceolata **     
                    |  |                 +--+                              
                    |  |                    +-- Lanceolaria_grayii[2] **      
                    +--+                                                   
                       |  +-------------------- Lepidodesma_languilati **     
                       |  |                                                
                       |  |              +----- Aculamprotula_tortuosa **     
                       |  |  +-----------+                                 
                       +--+  |           |  +-- Aculamprotula_tientsinensis **
                          |  |           +--+                              
                          |  |              +-- Aculamprotula_coreana **      
                          |  |                                             
                          +--+           +----- Unio_crassus **               
                             |  +--------+                                 
                             |  |        |  +-- Unio_delphinus **             
                             |  |        +--+                              
                             |  |           +-- Unio_pictorum **              
                             |  |                                          
                             +--+     +-------- Inversiunio_jokohamensis **   
                                |  +--+                                    
                                |  |  |  +----- Inversiunio_yanagawensis[2] **
                                |  |  +--+                                 
                                |  |     |  +-- Inversiunio_reinianus **      
                                |  |     +--+                              
                                +--+        +-- Inversiunio_yanagawensis[1] **
                                   |                                       
                                   |        +-- Tchangsinaia_piscicula **     
                                   |  +-----+                              
                                   |  |     +-- Schistodesmus_lampreyanus **  
                                   +--+                                    
                                      |  +----- Nodularia_douglasiae[2] **    
                                      +--+                                 
                                         |  +-- Nodularia_nipponensis **      
                                         +--+                              
                                            +-- Nodularia_douglasiae[1] **    

Dai et al., 2024 [Publ 4358], fig. 4

iT tT bi riS oT roT roS iD
50 50 48 3 2 2 1 86.0
                                         +-- Gibbosula_rochechouartii     
  +--------------------------------------+                                
  |                                      +-- Margaritifera_dahurica       
  |                                                                       
  |                                      +-- Quadrula_quadrula **            
  |                 +--------------------+                                
  |                 |                    +-- Uniomerus_tetralasmus **        
  |                 |                                                     
  |  +--------------+                    +-- Elliptio_complanata **          
  |  |              |  +-----------------+                                
  |  |              |  |                 +-- Pleurobema_oviforme **          
  |  |              +--+                                                  
  |  |                 |  +----------------- Amblema_plicata **              
--+  |                 |  |                                               
  |  |                 +--+  +-------------- Popenaias_popeii **             
  |  |                    |  |                                            
  |  |                    +--+  +----------- Toxolasma_parvum **             
  |  |                       |  |                                         
  |  |                       +--+        +-- Potamilus_leptodon **           
  |  |                          |  +-----+                                
  |  |                          |  |     +-- Potamilus_alatus **             
  |  |                          +--+                                      
  |  |                             |  +----- Leaunio_lienosus **             
  |  |                             +--+                                   
  |  |                                |  +-- Lampsilis_cardium **            
  |  |                                +--+                                
  +--+                                   +-- Venustaconcha_ellipsiformis **  
     |                                                                    
     |                                   +-- Chamberlainia_somsakpanhai **   
     |                    +--------------+                                
     |                    |              +-- Sinohyriopsis_cumingii **       
     |                    |                                               
     |                    |           +----- Potomida_littoralis **          
     |  +-----------------+  +--------+                                   
     |  |                 |  |        |  +-- Lamprotula_leaii **             
     |  |                 |  |        +--+                                
     |  |                 |  |           +-- Pronodularia_japanensis **      
     |  |                 +--+                                            
     |  |                    |           +-- Pseudodon_vondembuschianus **   
     |  |                    |  +--------+                                
     |  |                    |  |        +-- Pilsbryoconcha_exilis **        
     |  |                    +--+                                         
     |  |                       |  +-------- Microcondylaea_bonellii **      
     |  |                       +--+                                      
     |  |                          |  +----- Sinosolenaia_oleivora **        
     +--+                          +--+                                   
        |                             |  +-- Ptychorhynchus_pfisteri **      
        |                             +--+                                
        |                                +-- Parvasolenaia_rivularis **      
        |                                                                 
        |  +-------------------------------- Lepidodesma_languilati **       
        |  |                                                              
        |  |              +----------------- Lanceolaria_gladiola **         
        |  |              |                                               
        |  |  +-----------+  +-------------- Acuticosta_chinensis **         
        |  |  |           |  |                                            
        |  |  |           |  |        +----- Cristaria_plicata **            
        +--+  |           +--+  +-----+                                   
           |  |              |  |     |  +-- Anemina_arcaeformis **          
           |  |              |  |     +--+                                
           |  |              +--+        +-- Sinanodonta_woodiana **         
           |  |                 |                                         
           |  |                 |  +-------- Anodonta_anatina **             
           |  |                 +--+                                      
           +--+                    |  +----- Platynaias_compressa **         
              |                    +--+                                   
              |                       |  +-- Pyganodon_grandis **            
              |                       +--+                                
              |                          +-- Utterbackia_imbecillis **       
              |                                                           
              |                          +-- Aculamprotula_scripta **        
              |  +-----------------------+                                
              |  |                       +-- Aculamprotula_tientsinensis **  
              |  |                                                        
              |  |                       +-- Unio_crassus **                 
              +--+  +--------------------+                                
                 |  |                    +-- Unio_delphinus **               
                 |  |                                                     
                 |  |              +-------- Pseudocuneopsis_capitata **     
                 +--+  +-----------+                                      
                    |  |           |  +----- Pseudocuneopsis_yangshuoensis **
                    |  |           +--+                                   
                    |  |              |  +-- Pseudocuneopsis_yemaoi **       
                    |  |              +--+                                
                    +--+                 +-- Pseudocuneopsis_sichuanensis ** 
                       |                                                  
                       |                 +-- Nodularia_breviconcha **        
                       |  +--------------+                                
                       |  |              +-- Nodularia_douglasiae **         
                       |  |                                               
                       +--+              +-- Schistodesmus_lampreyanus **    
                          |  +-----------+                                
                          |  |           +-- Schistodesmus_spinosus **       
                          +--+                                            
                             |  +----------- Tchangsinaia_piscicula **       
                             +--+                                         
                                |  +-------- Cuneopsis_rufescens **          
                                +--+                                      
                                   |  +----- Cuneopsis_celtiformis **        
                                   +--+                                   
                                      |  +-- Cuneopsis_heudei **             
                                      +--+                                
                                         +-- Cuneopsis_demangei **           

Sano et al., 2020 [Publ 4138], fig. 3

iT tT bi riS oT roT roS iD
49 49 47 4 23 12 1 84.0
                       +----------------------- Cumberlandia_monodonta     
                       |                                                   
                       |                    +-- Pseudunio_auricularius     
  +--------------------+  +-----------------+                              
  |                    |  |                 +-- Pseudunio_marocanus        
  |                    |  |                                                
  |                    +--+                 +-- Gibbosula_laosensis        
  |                       |  +--------------+                              
  |                       |  |              +-- Gibbosula_rochechouartii   
  |                       |  |                                             
  |                       +--+              +-- Margaritifera_margaritifera
  |                          |  +-----------+                              
  |                          |  |           +-- Margaritifera_dahurica     
  |                          +--+                                          
  |                             |  +----------- Margaritifera_falcata      
  |                             +--+                                       
  |                                |  +-------- Margaritifera_middendorffi 
  |                                +--+                                    
--+                                   |  +----- Margaritifera_laevis       
  |                                   +--+                                 
  |                                      |  +-- Margaritifera_hembeli      
  |                                      +--+                              
  |                                         +-- Margaritifera_marrianae    
  |                                                                        
  |  +----------------------------------------- Scabies_crispata **           
  |  |                                                                     
  |  |                                   +----- Sinohyriopsis_cumingii[2] **  
  |  |                          +--------+                                 
  |  |                          |        |  +-- Sinohyriopsis_schlegelii **   
  |  |                          |        +--+                              
  |  |                          |           +-- Sinohyriopsis_cumingii[1] **  
  |  |  +-----------------------+                                          
  |  |  |                       |  +----------- Lampsilis_cardium **          
  +--+  |                       |  |                                       
     |  |                       |  |     +----- Lamprotula_leaii **           
     |  |                       +--+  +--+                                 
     |  |                          |  |  |  +-- Potomida_littoralis **        
     |  |                          |  |  +--+                              
     |  |                          +--+     +-- Pronodularia_japanensis **    
     |  |                             |                                    
     |  |                             |  +----- Inversidens_brandtii **       
     |  |                             +--+                                 
     |  |                                |  +-- Obovalis_omiensis **          
     +--+                                +--+                              
        |                                   +-- Ptychorhynchus_pfisteri **    
        |                                                                  
        |                                +----- Buldowskia_iwakawai **        
        |  +-----------------------------+                                 
        |  |                             |  +-- Anemina_arcaeformis **        
        |  |                             +--+                              
        |  |                                +-- Anemina_euscaphys **          
        |  |                                                               
        |  |                          +-------- Pletholophus_tenuis **        
        |  |                       +--+                                    
        |  |                       |  |  +----- Beringiana_fukuharai **       
        |  |                       |  +--+                                 
        +--+                       |     |  +-- Beringiana_japonica **        
           |  +--------------------+     +--+                              
           |  |                    |        +-- Beringiana_beringiana **      
           |  |                    |                                       
           |  |                    |  +-------- Sinanodonta_lucida **         
           |  |                    +--+                                    
           |  |                       |  +----- Sinanodonta_tumens **         
           |  |                       +--+                                 
           |  |                          |  +-- Sinanodonta_woodiana **       
           |  |                          +--+                              
           +--+                             +-- Sinanodonta_lauta **          
              |                                                            
              |  +----------------------------- Cristaria_plicata **          
              |  |                                                         
              |  |                          +-- Anodonta_anatina **           
              |  |                 +--------+                              
              |  |                 |        +-- Anodonta_cygnea **            
              |  |                 |                                       
              |  |  +--------------+     +----- Pyganodon_grandis **          
              +--+  |              |  +--+                                 
                 |  |              |  |  |  +-- Utterbackia_imbecillis **     
                 |  |              |  |  +--+                              
                 |  |              +--+     +-- Utterbackia_peninsularis **   
                 |  |                 |                                    
                 |  |                 |  +----- Alasmidonta_varicosa **       
                 |  |                 +--+                                 
                 +--+                    |  +-- Platynaias_compressa **       
                    |                    +--+                              
                    |                       +-- Prolasmidonta_heterodon **    
                    |                                                      
                    |                    +----- Lanceolaria_grayii[1] **      
                    |  +-----------------+                                 
                    |  |                 |  +-- Lanceolaria_lanceolata **     
                    |  |                 +--+                              
                    |  |                    +-- Lanceolaria_grayii[2] **      
                    +--+                                                   
                       |  +-------------------- Lepidodesma_languilati **     
                       |  |                                                
                       |  |              +----- Aculamprotula_tortuosa **     
                       |  |  +-----------+                                 
                       +--+  |           |  +-- Aculamprotula_tientsinensis **
                          |  |           +--+                              
                          |  |              +-- Aculamprotula_coreana **      
                          |  |                                             
                          +--+           +----- Unio_crassus **               
                             |  +--------+                                 
                             |  |        |  +-- Unio_delphinus **             
                             |  |        +--+                              
                             |  |           +-- Unio_pictorum **              
                             |  |                                          
                             +--+     +-------- Inversiunio_jokohamensis **   
                                |  +--+                                    
                                |  |  |  +----- Inversiunio_yanagawensis[2] **
                                |  |  +--+                                 
                                |  |     |  +-- Inversiunio_reinianus **      
                                |  |     +--+                              
                                +--+        +-- Inversiunio_yanagawensis[1] **
                                   |                                       
                                   |        +-- Nodularia_nipponensis **      
                                   |     +--+                              
                                   |     |  +-- Nodularia_douglasiae **       
                                   +-----+                                 
                                         |  +-- Schistodesmus_lampreyanus **  
                                         +--+                              
                                            +-- Tchangsinaia_piscicula **     

Sano et al., 2020 [Publ 4138], fig. 4

iT tT bi riS oT roT roS iD
49 49 47 4 23 12 1 83.0
                             +-------------------- Cumberlandia_monodonta     
                             |                                                
                             |                 +-- Pseudunio_auricularius     
  +--------------------------+  +--------------+                              
  |                          |  |              +-- Pseudunio_marocanus        
  |                          |  |                                             
  |                          +--+              +-- Gibbosula_rochechouartii   
  |                             |  +-----------+                              
  |                             |  |           +-- Gibbosula_laosensis        
  |                             |  |                                          
  |                             +--+           +-- Margaritifera_margaritifera
  |                                |  +--------+                              
  |                                |  |        +-- Margaritifera_dahurica     
  |                                +--+                                       
  |                                   |  +-------- Margaritifera_falcata      
  |                                   |  |                                    
  |                                   +--+     +-- Margaritifera_laevis       
  |                                      |  +--+                              
--+                                      |  |  +-- Margaritifera_middendorffi 
  |                                      +--+                                 
  |                                         |  +-- Margaritifera_hembeli      
  |                                         +--+                              
  |                                            +-- Margaritifera_marrianae    
  |                                                                           
  |  +-------------------------------------------- Scabies_crispata **           
  |  |                                                                        
  |  |                          +----------------- Lampsilis_cardium **          
  |  |                          |                                             
  |  |                          |           +----- Sinohyriopsis_cumingii[2] **  
  |  |  +-----------------------+  +--------+                                 
  |  |  |                       |  |        |  +-- Sinohyriopsis_schlegelii **   
  |  |  |                       |  |        +--+                              
  |  |  |                       +--+           +-- Sinohyriopsis_cumingii[1] **  
  +--+  |                          |                                          
     |  |                          |  +----------- Inversidens_brandtii **       
     |  |                          |  |                                       
     |  |                          +--+        +-- Obovalis_omiensis **          
     |  |                             |  +-----+                              
     |  |                             |  |     +-- Ptychorhynchus_pfisteri **    
     |  |                             +--+                                    
     |  |                                |  +----- Lamprotula_leaii **           
     |  |                                +--+                                 
     +--+                                   |  +-- Pronodularia_japanensis **    
        |                                   +--+                              
        |                                      +-- Potomida_littoralis **        
        |                                                                     
        |                                   +----- Buldowskia_iwakawai **        
        |  +--------------------------------+                                 
        |  |                                |  +-- Anemina_arcaeformis **        
        |  |                                +--+                              
        |  |                                   +-- Anemina_euscaphys **          
        |  |                                                                  
        |  |                             +-------- Sinanodonta_lucida **         
        |  |                          +--+                                    
        |  |                          |  |  +----- Sinanodonta_tumens **         
        |  |                          |  +--+                                 
        +--+                          |     |  +-- Sinanodonta_woodiana **       
           |  +-----------------------+     +--+                              
           |  |                       |        +-- Sinanodonta_lauta **          
           |  |                       |                                       
           |  |                       |  +-------- Pletholophus_tenuis **        
           |  |                       +--+                                    
           |  |                          |  +----- Beringiana_fukuharai **       
           |  |                          +--+                                 
           |  |                             |  +-- Beringiana_japonica **        
           |  |                             +--+                              
           +--+                                +-- Beringiana_beringiana **      
              |                                                               
              |  +-------------------------------- Cristaria_plicata **          
              |  |                                                            
              |  |                             +-- Anodonta_cygnea **            
              |  |                    +--------+                              
              |  |                    |        +-- Anodonta_anatina **           
              |  |                    |                                       
              |  |  +-----------------+     +----- Alasmidonta_varicosa **       
              +--+  |                 |  +--+                                 
                 |  |                 |  |  |  +-- Prolasmidonta_heterodon **    
                 |  |                 |  |  +--+                              
                 |  |                 +--+     +-- Platynaias_compressa **       
                 |  |                    |                                    
                 |  |                    |  +----- Pyganodon_grandis **          
                 |  |                    +--+                                 
                 +--+                       |  +-- Utterbackia_imbecillis **     
                    |                       +--+                              
                    |                          +-- Utterbackia_peninsularis **   
                    |                                                         
                    |  +-------------------------- Lepidodesma_languilati **     
                    |  |                                                      
                    |  |                    +----- Lanceolaria_grayii[1] **      
                    |  |  +-----------------+                                 
                    +--+  |                 |  +-- Lanceolaria_lanceolata **     
                       |  |                 +--+                              
                       |  |                    +-- Lanceolaria_grayii[2] **      
                       |  |                                                   
                       +--+                 +----- Aculamprotula_tortuosa **     
                          |  +--------------+                                 
                          |  |              |  +-- Aculamprotula_tientsinensis **
                          |  |              +--+                              
                          |  |                 +-- Aculamprotula_coreana **      
                          |  |                                                
                          +--+              +----- Unio_crassus **               
                             |  +-----------+                                 
                             |  |           |  +-- Unio_delphinus **             
                             |  |           +--+                              
                             |  |              +-- Unio_pictorum **              
                             +--+                                             
                                |              +-- Nodularia_douglasiae **       
                                |  +-----------+                              
                                |  |           +-- Nodularia_nipponensis **      
                                |  |                                          
                                +--+           +-- Schistodesmus_lampreyanus **  
                                   |  +--------+                              
                                   |  |        +-- Tchangsinaia_piscicula **     
                                   +--+                                       
                                      |  +-------- Inversiunio_jokohamensis **   
                                      +--+                                    
                                         |  +----- Inversiunio_yanagawensis[2] **
                                         +--+                                 
                                            |  +-- Inversiunio_reinianus **      
                                            +--+                              
                                               +-- Inversiunio_yanagawensis[1] **

Huang et al., 2019 [Publ 3843], fig. 3

iT tT bi riS oT roT roS iD
47 47 45 3 10 6 1 81.0
                       +-------------- Gibbosula_rochechouartii   
  +--------------------+                                          
  |                    |  +----------- Cumberlandia_monodonta     
  |                    +--+                                       
  |                       |  +-------- Pseudunio_marocanus        
  |                       +--+                                    
  |                          |  +----- Margaritifera_falcata      
  |                          +--+                                 
  |                             |  +-- Margaritifera_margaritifera
  |                             +--+                              
  |                                +-- Margaritifera_dahurica     
  |                                                               
  |                                +-- Sinohyriopsis_cumingii **     
  |                    +-----------+                              
  |                    |           +-- Sinohyriopsis_schlegelii **   
  |                    |                                          
  |                    |     +-------- Potomida_littoralis **        
  |           +--------+  +--+                                    
  |           |        |  |  |  +----- Pronodularia_japanensis **    
  |           |        |  |  +--+                                 
  |           |        |  |     |  +-- Lamprotula_caveata **         
  |           |        |  |     +--+                              
--+           |        +--+        +-- Lamprotula_leaii **           
  |           |           |                                       
  |           |           |        +-- Ptychorhynchus_pfisteri **    
  |           |           |     +--+                              
  |  +--------+           |     |  +-- Parvasolenaia_rivularis **    
  |  |        |           +-----+                                 
  |  |        |                 |  +-- Sinosolenaia_oleivora **      
  |  |        |                 +--+                              
  |  |        |                    +-- Sinosolenaia_carinata **      
  |  |        |                                                   
  |  |        |                    +-- Quadrula_quadrula **          
  |  |        |  +-----------------+                              
  |  |        |  |                 +-- Uniomerus_tetralasmus **      
  |  |        +--+                                                
  |  |           |  +----------------- Elliptio_complanata **        
  |  |           |  |                                             
  |  |           +--+  +-------------- Toxolasma_parvum **           
  |  |              |  |                                          
  |  |              +--+           +-- Potamilus_leptodon **         
  |  |                 |  +--------+                              
  |  |                 |  |        +-- Potamilus_alatus **           
  +--+                 +--+                                       
     |                    |  +-------- Leaunio_lienosus **           
     |                    +--+                                    
     |                       |  +----- Venustaconcha_ellipsiformis **
     |                       +--+                                 
     |                          |  +-- Lampsilis_cardium **          
     |                          +--+                              
     |                             +-- Lampsilis_ornata **           
     |                                                            
     |  +----------------------------- Lepidodesma_languilati **     
     |  |                                                         
     |  |                    +-------- Aculamprotula_tortuosa **     
     |  |              +-----+                                    
     |  |              |     |  +----- Aculamprotula_scripta **      
     |  |              |     +--+                                 
     |  |              |        |  +-- Aculamprotula_tientsinensis **
     |  |              |        +--+                              
     |  |  +-----------+           +-- Aculamprotula_coreana **      
     +--+  |           |                                          
        |  |           |        +----- Nodularia_douglasiae **       
        |  |           |  +-----+                                 
        |  |           |  |     |  +-- Tchangsinaia_piscicula **     
        |  |           |  |     +--+                              
        |  |           +--+        +-- Schistodesmus_lampreyanus **  
        |  |              |                                       
        |  |              |  +-------- Unio_tumidus **               
        |  |              +--+                                    
        +--+                 |  +----- Unio_crassus **               
           |                 +--+                                 
           |                    |  +-- Unio_delphinus **             
           |                    +--+                              
           |                       +-- Unio_pictorum **              
           |                                                      
           |                    +----- Lanceolaria_gladiola **       
           |  +-----------------+                                 
           |  |                 |  +-- Lanceolaria_lanceolata **     
           |  |                 +--+                              
           |  |                    +-- Lanceolaria_grayii **         
           +--+                                                   
              |  +-------------------- Acuticosta_chinensis **       
              |  |                                                
              |  |           +-------- Cristaria_plicata **          
              |  |           |                                    
              +--+  +--------+     +-- Sinanodonta_lucida **         
                 |  |        |  +--+                              
                 |  |        |  |  +-- Sinanodonta_woodiana **       
                 |  |        +--+                                 
                 +--+           |  +-- Anemina_arcaeformis **        
                    |           +--+                              
                    |              +-- Anemina_euscaphys **          
                    |                                             
                    |  +-------------- Anodonta_cygnea **            
                    +--+                                          
                       |  +----------- Anodonta_anatina **           
                       +--+                                       
                          |  +-------- Lasmigona_costata **          
                          +--+                                    
                             |  +----- Pyganodon_grandis **          
                             +--+                                 
                                |  +-- Utterbackia_imbecillis **     
                                +--+                              
                                   +-- Utterbackia_peninsularis **   

Bolotov et al., 2022 [Publ 4250], fig. 8

iT tT bi riS oT roT roS iD
47 49 42 2 2 2 1 94.0
                                +-- Gibbosula_laosensis          
                                |                                
                                +-- Margaritifera_dahurica       
                                |                                
                                +-- Sinosolenaia_iridinea **        
                                |                                
  +-----------------------------+-- Leguminaia_wheatleyi **         
  |                             |                                
  |                             +-- Gonidea_angulata **             
  |                             |                                
  |                             +-- Lamprotula_leaii **             
  |                             |                                
  |                             +-- Potomida_littoralis **          
  |                                                              
  |                             +-- Pseudodon_lenyanensis **        
  |                    +--------+                                
  |                    |        +-- Sundadontina_tanintharyiensis **
  |           +--------+                                         
  |           |        |  +-------- Indopseudodon_kayinensis **     
  |           |        +--+                                      
  |           |           |  +----- Indopseudodon_indawgyiensis **  
  |           |           +--+                                   
  |           |              |  +-- Indopseudodon_salwenianus **    
  |           |              +--+                                
  |  +--------+                 +-- Indopseudodon_bogani **         
--+  |        |                                                  
  |  |        |              +----- Trapezoideus_mitanensis **      
  |  |        |  +-----------+                                   
  |  |        |  |           |  +-- Trapezoideus_foliaceus **       
  |  |        |  |           +--+                                
  |  |        |  |              +-- Trapezoideus_lenya **           
  |  |        +--+                                               
  |  |           |              +-- Yaukthwa_elongatula **          
  |  |           |  +-----------+                                
  |  |           |  |           +-- Yaukthwa_baniensis **           
  |  |           |  |                                            
  |  |           +--+        +----- Yaukthwa_inlenensis **          
  |  |              |  +-----+                                   
  |  |              |  |     |  +-- Yaukthwa_paiensis **            
  |  |              |  |     +--+                                
  |  |              +--+        +-- Yaukthwa_avaensis **            
  |  |                 |                                         
  |  |                 |  +-------- Yaukthwa_panhai **              
  |  |                 +--+                                      
  +--+                    |  +----- Yaukthwa_rectangularis **       
     |                    +--+                                   
     |                       |  +-- Yaukthwa_nesemanni **           
     |                       +--+                                
     |                          +-- Yaukthwa_zayleymanensis **      
     |                                                           
     |                    +-------- Lamellidens_savadiensis **      
     |              +-----+                                      
     |              |     |  +----- Lamellidens_ferrugineus **      
     |              |     +--+                                   
     |              |        |  +-- Lamellidens_generosus **        
     |  +-----------+        +--+                                
     |  |           |           +-- Lamellidens_mainwaringi **      
     |  |           |                                            
     |  |           |  +----------- Trapezidens_exolescens **       
     |  |           +--+                                         
     |  |              |  +-------- Trapezidens_scutum **           
     |  |              +--+                                      
     |  |                 |  +----- Trapezidens_yeti **             
     |  |                 +--+                                   
     +--+                    |  +-- Trapezidens_angustior **        
        |                    +--+                                
        |                       +-- Trapezidens_dolichorhynchus **  
        |                                                        
        |  +----------------------- Parreysia_rakhinensis **        
        |  |                                                     
        |  |                    +-- Leoparreysia_whitteni **        
        |  |              +-----+                                
        |  |              |     +-- Leoparreysia_burmana **         
        +--+  +-----------+                                      
           |  |           |  +----- Leoparreysia_olivacea **        
           |  |           +--+                                   
           |  |              |  +-- Leoparreysia_canefrii **        
           |  |              +--+                                
           +--+                 +-- Leoparreysia_tavoyensis **      
              |                                                  
              |                 +-- Indonaia_andersoniana **        
              |  +--------------+                                
              |  |              +-- Indonaia_subclathrata **        
              +--+                                               
                 |  +-------------- Indochinella_pugio **           
                 |  |                                            
                 +--+  +----------- Radiatula_crispisulcata **      
                    |  |                                         
                    +--+        +-- Radiatula_chaudhurii **         
                       |  +-----+                                
                       |  |     +-- Radiatula_myitkyinae **         
                       +--+                                      
                          |  +----- Radiatula_mouhoti **            
                          +--+                                   
                             |  +-- Radiatula_chindwinensis **      
                             +--+                                
                                +-- Radiatula_myitthanensis **      

Zieritz et al., 2021 [Publ 4231], fig. 3-F, 5

iT tT bi riS oT roT roS iD
46 46 44 3 3 3 1 83.0
                             +----- Pseudunio_marocanus        
  +--------------------------+                                 
  |                          |  +-- Margaritifera_falcata      
  |                          +--+                              
  |                             +-- Margaritifera_dahurica     
  |                                                            
  |                    +----------- Quadrula_quadrula **          
  |                    |                                       
  |  +-----------------+  +-------- Toxolasma_parvum **           
  |  |                 |  |                                    
  |  |                 +--+     +-- Potamilus_alatus **           
  |  |                    |  +--+                              
  |  |                    |  |  +-- Potamilus_leptodon **         
--+  |                    +--+                                 
  |  |                       |  +-- Venustaconcha_ellipsiformis **
  |  |                       +--+                              
  |  |                          +-- Lampsilis_ornata **           
  |  |                                                         
  |  |                       +----- Chamberlainia_somsakpanhai ** 
  |  |        +--------------+                                 
  |  |        |              |  +-- Sinohyriopsis_cumingii **     
  |  |        |              +--+                              
  |  |        |                 +-- Sinohyriopsis_schlegelii **   
  |  |        |                                                
  |  |        |                 +-- Lens_contradens **            
  +--+  +-----+              +--+                              
     |  |     |              |  +-- Physunio_superbus **          
     |  |     |  +-----------+                                 
     |  |     |  |           |  +-- Hyriopsis_bialata **          
     |  |     |  |           +--+                              
     |  |     |  |              +-- Rectidens_sumatrensis **      
     |  |     |  |                                             
     |  |     +--+        +-------- Potomida_littoralis **        
     |  |        |  +-----+                                    
     |  |        |  |     |  +----- Pronodularia_japanensis **    
     |  |        |  |     +--+                                 
     |  |        |  |        |  +-- Lamprotula_caveata **         
     |  |        |  |        +--+                              
     |  |        +--+           +-- Lamprotula_leaii **           
     |  |           |                                          
     |  |           |           +-- Pilsbryoconcha_exilis **      
     |  |           |  +--------+                              
     +--+           |  |        +-- Pseudodon_vondembuschianus ** 
        |           +--+                                       
        |              |  +-------- Ptychorhynchus_pfisteri **    
        |              +--+                                    
        |                 |  +----- Microcondylaea_bonellii **    
        |                 +--+                                 
        |                    |  +-- Sinosolenaia_recognita **     
        |                    +--+                              
        |                       +-- Sinosolenaia_carinata **      
        |                                                      
        |  +----------------------- Lepidodesma_languilati **     
        |  |                                                   
        |  |                 +----- Aculamprotula_tortuosa **     
        |  |        +--------+                                 
        |  |        |        |  +-- Aculamprotula_tientsinensis **
        |  |        |        +--+                              
        |  |        |           +-- Aculamprotula_coreana **      
        |  |  +-----+                                          
        +--+  |     |        +----- Nodularia_douglasiae **       
           |  |     |  +-----+                                 
           |  |     |  |     |  +-- Schistodesmus_lampreyanus **  
           |  |     |  |     +--+                              
           |  |     +--+        +-- Tchangsinaia_piscicula **     
           |  |        |                                       
           |  |        |  +-------- Unio_tumidus **               
           |  |        +--+                                    
           |  |           |  +----- Unio_crassus **               
           +--+           +--+                                 
              |              |  +-- Unio_pictorum **              
              |              +--+                              
              |                 +-- Unio_delphinus **             
              |                                                
              |                 +-- Lanceolaria_grayii **         
              |  +--------------+                              
              |  |              +-- Lanceolaria_lanceolata **     
              |  |                                             
              |  |     +----------- Anodonta_anatina **           
              +--+  +--+                                       
                 |  |  |  +-------- Platynaias_compressa **       
                 |  |  +--+                                    
                 |  |     |  +----- Pyganodon_grandis **          
                 |  |     +--+                                 
                 +--+        |  +-- Utterbackia_imbecillis **     
                    |        +--+                              
                    |           +-- Utterbackia_peninsularis **   
                    |                                          
                    |     +-------- Cristaria_plicata **          
                    |     |                                    
                    +-----+     +-- Sinanodonta_lucida **         
                          |  +--+                              
                          |  |  +-- Sinanodonta_woodiana **       
                          +--+                                 
                             |  +-- Anemina_arcaeformis **        
                             +--+                              
                                +-- Anemina_euscaphys **          

Burlakova et al., 2012 [Publ 3486], fig. 1

iT tT bi riS oT roT roS iD
44 44 42 1 0 0 0 81.0
                                               +-- Amblema_elliottii **       
  +--------------------------------------------+                           
  |                                            +-- Amblema_plicata **         
  |                                                                        
--+                                            +-- Lampsilis_ovata **         
  |  +-----------------------------------------+                           
  |  |                                         +-- Plectomerus_dombeyanus **  
  |  |                                                                     
  +--+  +----------------------------------------- Popenaias_popeii **        
     |  |                                                                  
     |  |  +-------------------------------------- Cyrtonaias_tampicoensis ** 
     +--+  |                                                               
        |  |  +----------------------------------- Parvaspina_collina **      
        |  |  |                                                            
        +--+  |                             +----- Plethobasus_cyphyus **     
           |  |  +--------------------------+                              
           |  |  |                          |  +-- Elliptio_arca **           
           |  |  |                          +--+                           
           |  |  |                             +-- Elliptio_crassidens **     
           +--+  |                                                         
              |  |           +-------------------- Fusconaia_subrotunda **    
              |  |           |                                             
              |  |           |                 +-- Fusconaia_cor **           
              |  |  +--------+  +--------------+                           
              |  |  |        |  |              +-- Fusconaia_cuneolus **      
              |  |  |        |  |                                          
              +--+  |        +--+  +-------------- Fusconaia_ozarkensis **    
                 |  |           |  |                                       
                 |  |           |  |     +-------- Fusconaia_askewi[3] **     
                 |  |           |  |  +--+                                 
                 |  |           +--+  |  |  +----- Fusconaia_askewi[2] **     
                 |  |              |  |  +--+                              
                 |  |              |  |     |  +-- Fusconaia_lananensis **    
                 |  |              |  |     +--+                           
                 |  |              +--+        +-- Fusconaia_askewi[1] **     
                 |  |                 |                                    
                 |  |                 |        +-- Fusconaia_burkei **        
                 |  |                 |  +-----+                           
                 +--+                 |  |     +-- Fusconaia_escambia **      
                    |                 +--+                                 
                    |                    |  +----- Fusconaia_masoni **        
                    |                    +--+                              
                    |                       |  +-- Fusconaia_flava **         
                    |                       +--+                           
                    |                          +-- Fusconaia_cerina **        
                    |                                                      
                    |                          +-- Eurynia_dilatata **        
                    |  +-----------------------+                           
                    |  |                       +-- Hemistena_lata **          
                    |  |                                                   
                    |  |                    +----- Pleuronaia_gibber **       
                    |  |              +-----+                              
                    |  |              |     |  +-- Pleuronaia_dolabelloides **
                    +--+              |     +--+                           
                       |  +-----------+        +-- Pleuronaia_barnesiana **   
                       |  |           |                                    
                       |  |           |  +-------- Pleurobema_riddellii **    
                       |  |           +--+                                 
                       |  |              |  +----- Pleurobema_sintoxia **     
                       |  |              +--+                              
                       +--+                 |  +-- Pleurobema_rubrum **       
                          |                 +--+                           
                          |                    +-- Pleurobema_cordatum **     
                          |                                                
                          |  +-------------------- Pleurobema_stabile **      
                          |  |                                             
                          +--+  +----------------- Pleurobema_pyriforme **    
                             |  |                                          
                             |  |           +----- Pleurobema_georgianum **   
                             +--+  +--------+                              
                                |  |        |  +-- Pleurobema_hanleyianum **  
                                |  |        +--+                           
                                |  |           +-- Pleurobema_decisum **      
                                +--+                                       
                                   |           +-- Pleurobema_clava **        
                                   |  +--------+                           
                                   |  |        +-- Pleurobema_oviforme **     
                                   +--+                                    
                                      |  +-------- Pleurobema_beadleianum **  
                                      |  |                                 
                                      +--+     +-- Pleurobema_rubellum **     
                                         |  +--+                           
                                         |  |  +-- Pleurobema_strodeanum **   
                                         +--+                              
                                            |  +-- Pleurobema_perovatum **    
                                            +--+                           
                                               +-- Pleurobema_taitianum **    

Inoue et al., 2020 [Publ 3941], fig. 3

iT tT bi riS oT roT roS iD
44 44 42 2 0 0 0 74.0
  +----------------------------------------------- Arcidens_confragosus **    
  |                                                                        
  |                                            +-- Pustulosa_petrina **       
  |  +-----------------------------------------+                           
  |  |                                         +-- Quadrula_quadrula **       
--+  |                                                                     
  |  |                             +-------------- Parvaspina_collina **      
  |  |                             |                                       
  |  |                             |           +-- Eurynia_dilatata **        
  |  |  +--------------------------+  +--------+                           
  |  |  |                          |  |        +-- Hemistena_lata **          
  +--+  |                          |  |                                    
     |  |                          +--+        +-- Pleurobema_clava **        
     |  |                             |  +-----+                           
     |  |                             |  |     +-- Pleurobema_sintoxia **     
     |  |                             +--+                                 
     |  |                                |  +----- Pleuronaia_dolabelloides **
     |  |                                +--+                              
     |  |                                   |  +-- Fusconaia_flava **         
     +--+                                   +--+                           
        |                                      +-- Fusconaia_subrotunda **    
        |                                                                  
        |                                      +-- Amblema_plicata **         
        |                                   +--+                           
        |                                   |  +-- Amblema_elliottii **       
        |  +--------------------------------+                              
        |  |                                |  +-- Reginaia_ebenus **         
        |  |                                +--+                           
        |  |                                   +-- Reginaia_rotulata **       
        |  |                                                               
        +--+  +----------------------------------- Toxolasma_lividum **       
           |  |                                                            
           |  |                          +-------- Potamilus_fragilis **      
           |  |                          |                                 
           |  |  +-----------------------+     +-- Potamilus_ohiensis **      
           +--+  |                       |  +--+                           
              |  |                       |  |  +-- Potamilus_amphichaenus **  
              |  |                       +--+                              
              |  |                          |  +-- Potamilus_metnecktayi **   
              |  |                          +--+                           
              |  |                             +-- Potamilus_purpuratus **    
              +--+                                                         
                 |                       +-------- Ellipsaria_lineolata **    
                 |  +--------------------+                                 
                 |  |                    |  +----- Truncilla_macrodon **      
                 |  |                    +--+                              
                 |  |                       |  +-- Truncilla_donaciformis **  
                 |  |                       +--+                           
                 |  |                          +-- Truncilla_cognata **       
                 +--+                                                      
                    |  +-------------------------- Leaunio_lienosus **        
                    |  |                                                   
                    |  |     +-------------------- Lampsilis_bracteata **     
                    |  |     |                                             
                    |  |  +--+  +----------------- Lampsilis_fasciola **      
                    |  |  |  |  |                                          
                    +--+  |  +--+  +-------------- Lampsilis_ornata **        
                       |  |     |  |                                       
                       |  |     +--+           +-- Lampsilis_ovata **         
                       |  |        |  +--------+                           
                       |  |        |  |        +-- Lampsilis_cariosa **       
                       |  |        +--+                                    
                       |  |           |  +-------- Lampsilis_cardium[3] **    
                       +--+           +--+                                 
                          |              |  +----- Lampsilis_cardium[2] **    
                          |              +--+                              
                          |                 |  +-- Lampsilis_satura **        
                          |                 +--+                           
                          |                    +-- Lampsilis_cardium[1] **    
                          |                                                
                          |                    +-- Lampsilis_higginsii **     
                          |     +--------------+                           
                          |     |              +-- Ortmanniana_ligamentina ** 
                          +-----+                                          
                                |  +-------------- Lampsilis_teres **         
                                |  |                                       
                                +--+           +-- Lampsilis_radiata **       
                                   |  +--------+                           
                                   |  |        +-- Lampsilis_siliquoidea **   
                                   +--+                                    
                                      |  +-------- Lampsilis_bergmanni **     
                                      +--+                                 
                                         |  +----- Lampsilis_hydiana **       
                                         +--+                              
                                            |  +-- Lampsilis_virescens **     
                                            +--+                           
                                               +-- Lampsilis_straminea **     

Sano et al., 2020 [Publ 4138], fig. 1

iT tT bi riS oT roT roS iD
44 44 13 2 4 3 1 75.0
              +----- Gibbosula_rochechouartii   
  +-----------+                                 
  |           |  +-- Margaritifera_middendorffi 
  |           +--+                              
  |              +-- Margaritifera_laevis       
  |                                             
  |        +-------- Sinosolenaia_iridinea **      
  |        |                                    
  |     +--+  +----- Inversidens_brandtii **       
  |     |  |  |                                 
  |     |  +--+  +-- Obovalis_omiensis[1] **       
  |     |     |  |                              
  |     |     +--+-- Obovalis_omiensis[2] **       
  |  +--+        |                              
  |  |  |        +-- Ptychorhynchus_pfisteri **    
  |  |  |                                       
  |  |  |        +-- Lamprotula_caveata **         
--+  |  |     +--+                              
  |  |  |     |  +-- Pronodularia_japanensis **    
  |  |  +-----+                                 
  |  |        |  +-- Sinohyriopsis_schlegelii[1] **
  |  |        |  |                              
  |  |        +--+-- Sinohyriopsis_schlegelii[2] **
  |  |           |                              
  |  |           +-- Sinohyriopsis_cumingii **     
  |  |                                          
  |  |     +-------- Lanceolaria_grayii[1] **      
  |  |     |                                    
  |  |     +-------- Lanceolaria_grayii[2] **      
  |  |     |                                    
  |  |     +-------- Cristaria_plicata[1] **       
  |  |     |                                    
  |  |     +-------- Cristaria_plicata[2] **       
  +--+     |                                    
     |     +-------- Sinanodonta_lauta **          
     |     |                                    
     |     +-------- Beringiana_japonica **        
     |     |                                    
     |     +-------- Beringiana_fukuharai[1] **    
     |     |                                    
     |     +-------- Beringiana_beringiana **      
     |     |                                    
     |     +-------- Beringiana_fukuharai[2] **    
     |     |                                    
     |     +-------- Beringiana_fukuharai[3] **    
     |     |                                    
     |     +-------- Buldowskia_iwakawai[1] **     
     |     |                                    
     |     +-------- Buldowskia_iwakawai[2] **     
     |     |                                    
     |     +-------- Sinanodonta_tumens[1] **      
     |     |                                    
     |     +-------- Sinanodonta_tumens[2] **      
     |     |                                    
     +-----+-------- Pletholophus_tenuis **        
           |                                    
           +-------- Anemina_arcaeformis **        
           |                                    
           +-------- Lanceolaria_lanceolata **     
           |                                    
           +-------- Cristaria_plicata[3] **       
           |                                    
           +-------- Lanceolaria_gladiola **       
           |                                    
           +-------- Sinanodonta_angula **         
           |                                    
           +-------- Lepidodesma_languilati **     
           |                                    
           |     +-- Aculamprotula_tortuosa **     
           +-----+                              
           |     +-- Acuticosta_chinensis **       
           |                                    
           |  +----- Cuneopsis_celtiformis **      
           |  |                                 
           +--+  +-- Nodularia_douglasiae[1] **    
           |  |  |                              
           |  +--+-- Nodularia_douglasiae[2] **    
           |     |                              
           |     +-- Nodularia_nipponensis **      
           |                                    
           |  +----- Schistodesmus_lampreyanus **  
           |  |                                 
           |  |  +-- Inversiunio_reinianus[1] **   
           +--+  |                              
              |  +-- Inversiunio_reinianus[2] **   
              |  |                              
              |  +-- Inversiunio_jokohamensis[1] **
              +--+                              
                 +-- Inversiunio_jokohamensis[2] **
                 |                              
                 +-- Inversiunio_jokohamensis[3] **
                 |                              
                 +-- Inversiunio_jokohamensis[4] **

Whelan et al., 2023 [Publ 4287], fig. 2

iT tT bi riS oT roT roS iD
44 44 42 1 0 0 0 83.0
                                +-------- Anemina_arcaeformis **             
  +-----------------------------+                                         
  |                             |  +----- Sinanodonta_lucida **              
  |                             +--+                                      
  |                                |  +-- Sinanodonta_woodiana **            
  |                                +--+                                   
--+                                   +-- Sinanodonta_tumens **              
  |                                                                       
  |  +----------------------------------- Anodonta_cygnea **                 
  |  |                                                                    
  |  |                                +-- Anodonta_nuttalliana **            
  |  |  +-----------------------------+                                   
  +--+  |                             +-- Pseudanodonta_complanata **        
     |  |                                                                 
     |  |  +----------------------------- Anodonta_anatina **                
     |  |  |                                                              
     |  |  |                       +----- Utterbackia_peninsularis[2] **     
     +--+  |                    +--+                                      
        |  |                    |  |  +-- Utterbackia_peninsularis[1] **     
        |  |                    |  +--+                                   
        |  |  +-----------------+     +-- Utterbackia_imbecillis **          
        |  |  |                 |                                         
        |  |  |                 |  +----- Pyganodon_grandis[2] **            
        +--+  |                 +--+                                      
           |  |                    |  +-- Pyganodon_grandis[1] **            
           |  |                    +--+                                   
           |  |                       +-- Pyganodon_lacustris **             
           |  |                                                           
           |  |                       +-- Utterbackiana_couperiana **        
           |  |                    +--+                                   
           |  |                    |  +-- Utterbackiana_heardi **            
           |  |              +-----+                                      
           |  |              |     |  +-- Utterbackiana_suborbiculata **     
           +--+              |     +--+                                   
              |              |        +-- Utterbackiana_hartfieldorum **     
              |  +-----------+                                            
              |  |           |     +----- Lasmigona_costata **               
              |  |           |  +--+                                      
              |  |           |  |  |  +-- Pseudodontoideus_subvexus **       
              |  |           |  |  +--+                                   
              |  |           +--+     +-- Pseudodontoideus_connasaugaensis **
              |  |              |                                         
              |  |              |  +----- Lasmigona_complanata **            
              |  |              +--+                                      
              |  |                 |  +-- Arcidens_confragosus **            
              |  |                 +--+                                   
              |  |                    +-- Arcidens_wheeleri **               
              +--+                                                        
                 |     +----------------- Pressodonta_viridis **             
                 |     |                                                  
                 |  +--+              +-- Anodontoides_denigratus **         
                 |  |  |  +-----------+                                   
                 |  |  |  |           +-- Anodontoides_ferussacianus **      
                 |  |  +--+                                               
                 |  |     |  +----------- Strophitus_undulatus **            
                 |  |     +--+                                            
                 |  |        |  +-------- Strophitus_pascagoulaensis[2] **   
                 |  |        +--+                                         
                 |  |           |  +----- Strophitus_radiatus **             
                 +--+           +--+                                      
                    |              |  +-- Strophitus_pascagoulaensis[1] **   
                    |              +--+                                   
                    |                 +-- Strophitus_williamsi **            
                    |                                                     
                    |              +----- Prolasmidonta_heterodon **         
                    |  +-----------+                                      
                    |  |           |  +-- Platynaias_subviridis **           
                    |  |           +--+                                   
                    |  |              +-- Platynaias_compressa **            
                    +--+                                                  
                       |              +-- Alasmidonta_holstonia **           
                       |  +-----------+                                   
                       |  |           +-- Alasminota_etowaensis **           
                       |  |                                               
                       +--+           +-- Alasmidonta_raveneliana **         
                          |  +--------+                                   
                          |  |        +-- Alasmidonta_marginata **           
                          |  |                                            
                          +--+        +-- Alasmidonta_varicosa **            
                             |  +-----+                                   
                             |  |     +-- Alasmidonta_uwharriensis **        
                             +--+                                         
                                |  +----- Alasmidonta_undulata **            
                                +--+                                      
                                   |  +-- Alasmidonta_triangulata **         
                                   +--+                                   
                                      +-- Alasmidonta_arcula **              

Campbell & Lydeard, 2012 [Publ 3308], fig. 1

iT tT bi riS oT roT roS iD
43 43 41 1 0 0 0 79.0
                                      +----------- Quadrula_quadrula **       
                                      |                                    
  +-----------------------------------+        +-- Pustulosa_infucata **      
  |                                   |  +-----+                           
  |                                   |  |     +-- Pustulosa_kleiniana **     
  |                                   +--+                                 
  |                                      |  +----- Cyclonaias_tuberculata **  
  |                                      +--+                              
  |                                         |  +-- Pustulosa_succissa **      
  |                                         +--+                           
  |                                            +-- Pustulosa_pustulosa **     
  |                                                                        
  |                                         +----- Popenaias_popeii **        
  |                             +-----------+                              
--+                             |           |  +-- Amblema_elliottii **       
  |                             |           +--+                           
  |                             |              +-- Amblema_plicata **         
  |  +--------------------------+                                          
  |  |                          |           +----- Cyrtonaias_tampicoensis ** 
  |  |                          |  +--------+                              
  |  |                          |  |        |  +-- Reginaia_ebenus **         
  |  |                          |  |        +--+                           
  |  |                          +--+           +-- Reginaia_rotulata **       
  |  |                             |                                       
  |  |                             |  +----------- Plectomerus_dombeyanus **  
  |  |                             +--+                                    
  |  |                                |  +-------- Lemiox_rimosus **          
  +--+                                +--+                                 
     |                                   |  +----- Lampsilis_ovata **         
     |                                   +--+                              
     |                                      |  +-- Obovaria_unicolor **       
     |                                      +--+                           
     |                                         +-- Obovaria_olivaria **       
     |                                                                     
     |  +----------------------------------------- Elliptoideus_sloatianus ** 
     |  |                                                                  
     |  |  +-------------------------------------- Pleurobema_stabile **      
     +--+  |                                                               
        |  |                                +----- Pleurobema_pyriforme **    
        |  |  +-----------------------------+                              
        +--+  |                             |  +-- Pleurobema_clava **        
           |  |                             +--+                           
           |  |                                +-- Pleurobema_decisum **      
           |  |                                                            
           +--+                                +-- Pleurobema_cordatum **     
              |  +-----------------------------+                           
              |  |                             +-- Pleurobema_sintoxia **     
              |  |                                                         
              |  |                             +-- Eurynia_dilatata **        
              |  |                       +-----+                           
              +--+                       |     +-- Hemistena_lata **          
                 |  +--------------------+                                 
                 |  |                    |  +----- Plethobasus_cyphyus **     
                 |  |                    +--+                              
                 |  |                       |  +-- Elliptio_arca **           
                 |  |                       +--+                           
                 +--+                          +-- Elliptio_crassidens **     
                    |                                                      
                    |                       +----- Pleuronaia_gibber **       
                    |  +--------------------+                              
                    |  |                    |  +-- Pleuronaia_barnesiana **   
                    |  |                    +--+                           
                    +--+                       +-- Pleuronaia_dolabelloides **
                       |                                                   
                       |  +----------------------- Parvaspina_collina **      
                       |  |                                                
                       +--+  +-------------------- Fusconaia_subrotunda **    
                          |  |                                             
                          +--+                 +-- Fusconaia_cor **           
                             |  +--------------+                           
                             |  |              +-- Fusconaia_cuneolus **      
                             +--+                                          
                                |  +-------------- Fusconaia_ozarkensis **    
                                |  |                                       
                                +--+           +-- Fusconaia_burkei **        
                                   |  +--------+                           
                                   |  |        +-- Fusconaia_escambia **      
                                   +--+                                    
                                      |  +-------- Fusconaia_masoni **        
                                      +--+                                 
                                         |  +----- Fusconaia_cerina[2] **     
                                         +--+                              
                                            |  +-- Fusconaia_flava **         
                                            +--+                           
                                               +-- Fusconaia_cerina[1] **     

Campbell & Lydeard, 2012 [Publ 3308], fig. 2

iT tT bi riS oT roT roS iD
43 43 36 1 0 0 0 77.0
                       +----------- Quadrula_quadrula **       
                       |                                    
  +--------------------+        +-- Pustulosa_infucata **      
  |                    |  +-----+                           
  |                    |  |     +-- Pustulosa_kleiniana **     
  |                    +--+                                 
  |                       |  +----- Cyclonaias_tuberculata **  
  |                       +--+                              
  |                          |  +-- Pustulosa_succissa **      
  |                          +--+                           
  |                             +-- Pustulosa_pustulosa **     
  |                                                         
  |                             +-- Reginaia_ebenus **         
  |                          +--+                           
  |                          |  +-- Reginaia_rotulata **       
--+              +-----------+                              
  |              |           |  +-- Amblema_elliottii **       
  |              |           +--+                           
  |  +-----------+              +-- Amblema_plicata **         
  |  |           |                                          
  |  |           |  +-------------- Popenaias_popeii **        
  |  |           |  |                                       
  |  |           +--+  +----------- Cyrtonaias_tampicoensis ** 
  |  |              |  |                                    
  |  |              +--+----------- Plectomerus_dombeyanus **  
  |  |                 |                                    
  |  |                 |  +-------- Lemiox_rimosus **          
  |  |                 +--+                                 
  |  |                    |  +----- Lampsilis_ovata **         
  +--+                    +--+                              
     |                       |  +-- Obovaria_unicolor **       
     |                       +--+                           
     |                          +-- Obovaria_olivaria **       
     |                                                      
     |  +-------------------------- Elliptoideus_sloatianus ** 
     |  |                                                   
     |  |  +----------------------- Hemistena_lata **          
     |  |  |                                                
     |  |  +----------------------- Plethobasus_cyphyus **     
     |  |  |                                                
     +--+  |                    +-- Elliptio_arca **           
        |  +--------------------+                           
        |  |                    +-- Elliptio_crassidens **     
        |  |                                                
        |  |                    +-- Eurynia_dilatata **        
        |  +--------------------+                           
        |  |                    +-- Parvaspina_collina **      
        +--+                                                
           |                 +----- Pleuronaia_gibber **       
           |  +--------------+                              
           |  |              |  +-- Pleuronaia_barnesiana **   
           |  |              +--+                           
           |  |                 +-- Pleuronaia_dolabelloides **
           |  |                                             
           |  |                 +-- Pleurobema_cordatum **     
           |  |        +--------+                           
           |  |        |        +-- Pleurobema_sintoxia **     
           |  +--------+                                    
           |  |        |  +-------- Pleurobema_stabile **      
           +--+        +--+                                 
              |           |  +----- Pleurobema_pyriforme **    
              |           +--+                              
              |              |  +-- Pleurobema_clava **        
              |              +--+                           
              |                 +-- Pleurobema_decisum **      
              |                                             
              |              +----- Fusconaia_subrotunda **    
              |  +-----------+                              
              |  |           |  +-- Fusconaia_cor **           
              |  |           +--+                           
              +--+              +-- Fusconaia_cuneolus **      
                 |                                          
                 |  +-------------- Fusconaia_ozarkensis **    
                 |  |                                       
                 +--+           +-- Fusconaia_burkei **        
                    |  +--------+                           
                    |  |        +-- Fusconaia_escambia **      
                    +--+                                    
                       |  +-------- Fusconaia_masoni **        
                       +--+                                 
                          |  +----- Fusconaia_cerina[2] **     
                          +--+                              
                             |  +-- Fusconaia_flava **         
                             +--+                           
                                +-- Fusconaia_cerina[1] **     

Campbell & Lydeard, 2012 [Publ 3308], fig. 4

iT tT bi riS oT roT roS iD
43 43 40 1 0 0 0 77.0
                                         +----------- Quadrula_quadrula **       
                                         |                                    
  +--------------------------------------+        +-- Pustulosa_infucata **      
  |                                      |  +-----+                           
  |                                      |  |     +-- Pustulosa_kleiniana **     
  |                                      +--+                                 
  |                                         |  +----- Cyclonaias_tuberculata **  
  |                                         +--+                              
  |                                            |  +-- Pustulosa_succissa **      
  |                                            +--+                           
  |                                               +-- Pustulosa_pustulosa **     
  |                                                                           
  |                                            +----- Popenaias_popeii **        
  |                                   +--------+                              
--+                                   |        |  +-- Amblema_elliottii **       
  |                                   |        +--+                           
  |                                   |           +-- Amblema_plicata **         
  |  +--------------------------------+                                       
  |  |                                |  +----------- Plectomerus_dombeyanus **  
  |  |                                |  |                                    
  |  |                                |  |     +----- Cyrtonaias_tampicoensis ** 
  |  |                                |  +-----+                              
  |  |                                +--+     |  +-- Reginaia_ebenus **         
  |  |                                   |     +--+                           
  |  |                                   |        +-- Reginaia_rotulata **       
  |  |                                   |                                    
  +--+                                   |  +-------- Lemiox_rimosus **          
     |                                   +--+                                 
     |                                      |  +----- Lampsilis_ovata **         
     |                                      +--+                              
     |                                         |  +-- Obovaria_unicolor **       
     |                                         +--+                           
     |                                            +-- Obovaria_olivaria **       
     |                                                                        
     |  +-------------------------------------------- Elliptoideus_sloatianus ** 
     |  |                                                                     
     +--+  +----------------------------------------- Pleurobema_stabile **      
        |  |                                                                  
        +--+  +-------------------------------------- Hemistena_lata **          
           |  |                                                               
           +--+  +----------------------------------- Eurynia_dilatata **        
              |  |                                                            
              |  |                                +-- Pleurobema_cordatum **     
              +--+  +-----------------------------+                           
                 |  |                             +-- Pleurobema_sintoxia **     
                 |  |                                                         
                 |  |                          +----- Pleurobema_clava **        
                 +--+  +-----------------------+                              
                    |  |                       |  +-- Pleurobema_decisum **      
                    |  |                       +--+                           
                    |  |                          +-- Pleurobema_pyriforme **    
                    |  |                                                      
                    +--+                    +-------- Parvaspina_collina **      
                       |  +-----------------+                                 
                       |  |                 |  +----- Pleuronaia_gibber **       
                       |  |                 +--+                              
                       |  |                    |  +-- Pleuronaia_barnesiana **   
                       |  |                    +--+                           
                       +--+                       +-- Pleuronaia_dolabelloides **
                          |                                                   
                          |                    +----- Plethobasus_cyphyus **     
                          |  +-----------------+                              
                          |  |                 |  +-- Elliptio_arca **           
                          |  |                 +--+                           
                          +--+                    +-- Elliptio_crassidens **     
                             |                                                
                             |  +-------------------- Fusconaia_subrotunda **    
                             |  |                                             
                             +--+                 +-- Fusconaia_cor **           
                                |  +--------------+                           
                                |  |              +-- Fusconaia_cuneolus **      
                                +--+                                          
                                   |  +-------------- Fusconaia_ozarkensis **    
                                   |  |                                       
                                   +--+           +-- Fusconaia_burkei **        
                                      |  +--------+                           
                                      |  |        +-- Fusconaia_escambia **      
                                      +--+                                    
                                         |  +-------- Fusconaia_masoni **        
                                         +--+                                 
                                            |  +----- Fusconaia_cerina[2] **     
                                            +--+                              
                                               |  +-- Fusconaia_flava **         
                                               +--+                           
                                                  +-- Fusconaia_cerina[1] **     

Burlakova et al., 2012 [Publ 3486], fig. 3

iT tT bi riS oT roT roS iD
43 43 26 1 0 0 0 75.0
  +----------------------------------- Plectomerus_dombeyanus **  
  |                                                            
  |  +-------------------------------- Hemistena_lata **          
  |  |                                                         
--+  +-------------------------------- Pleurobema_stabile **      
  |  |                                                         
  |  |  +----------------------------- Eurynia_dilatata **        
  |  |  |                                                      
  +--+  |                    +-------- Pleurobema_riddellii **    
     |  |                    |                                 
     |  |  +-----------------+  +----- Pleurobema_sintoxia **     
     |  |  |                 |  |                              
     |  |  |                 +--+----- Pleurobema_rubrum[2] **    
     |  |  |                    |                              
     +--+  |                    |  +-- Pleurobema_cordatum **     
        |  |                    +--+                           
        |  |                       +-- Pleurobema_rubrum[1] **    
        |  |                                                   
        |  |  +----------------------- Parvaspina_collina **      
        |  |  |                                                
        |  |  |                 +----- Pleuronaia_gibber **       
        |  |  +-----------------+                              
        |  |  |                 |  +-- Pleuronaia_dolabelloides **
        +--+  |                 +--+                           
           |  |                    +-- Pleuronaia_barnesiana **   
           |  |                                                
           |  |                 +----- Plethobasus_cyphyus **     
           |  +-----------------+                              
           |  |                 |  +-- Elliptio_arca **           
           |  |                 +--+                           
           |  |                    +-- Elliptio_crassidens **     
           |  |                                                
           |  |                    +-- Pleurobema_clava **        
           |  |           +--------+                           
           |  |           |        +-- Pleurobema_oviforme **     
           |  |           |                                    
           |  |           |  +-------- Pleurobema_decisum **      
           +--+-----------+  |                                 
              |           |  +-------- Pleurobema_pyriforme **    
              |           |  |                                 
              |           |  +-------- Pleurobema_hanleyianum **  
              |           +--+                                 
              |              +-------- Pleurobema_georgianum **   
              |              |                                 
              |              |  +----- Pleurobema_rubellum **     
              |              |  |                              
              |              |  +----- Pleurobema_beadleianum **  
              |              +--+                              
              |                 +----- Pleurobema_strodeanum **   
              |                 |                              
              |                 |  +-- Pleurobema_perovatum **    
              |                 +--+                           
              |                    +-- Pleurobema_taitianum **    
              |                                                
              |  +-------------------- Fusconaia_subrotunda **    
              |  |                                             
              |  |                 +-- Fusconaia_cor **           
              +--+  +--------------+                           
                 |  |              +-- Fusconaia_cuneolus **      
                 |  |                                          
                 |  |  +-------------- Fusconaia_ozarkensis **    
                 +--+  |                                       
                    |  |     +-------- Fusconaia_masoni **        
                    |  |     |                                 
                    |  |     |     +-- Fusconaia_burkei **        
                    |  |  +--+-----+                           
                    +--+  |  |     +-- Fusconaia_escambia **      
                       |  |  |                                 
                       |  |  |  +----- Fusconaia_cerina **        
                       |  |  +--+                              
                       |  |     |  +-- Fusconaia_askewi[1] **     
                       |  |     +--+                           
                       +--+        +-- Fusconaia_flava **         
                          |                                    
                          |        +-- Fusconaia_askewi[2] **     
                          |        |                           
                          |        +-- Fusconaia_askewi[3] **     
                          |        |                           
                          |        +-- Fusconaia_askewi[4] **     
                          +--------+                           
                                   +-- Fusconaia_lananensis[1] ** 
                                   |                           
                                   +-- Fusconaia_lananensis[2] ** 
                                   |                           
                                   +-- Fusconaia_askewi[5] **     

Inoue et al., 2020 [Publ 3941], fig. 4

iT tT bi riS oT roT roS iD
43 43 41 2 0 0 0 73.0
  +-------------------------------------------------- Arcidens_confragosus **    
  |                                                                           
  |                                               +-- Pustulosa_petrina **       
  |  +--------------------------------------------+                           
--+  |                                            +-- Quadrula_quadrula **       
  |  |                                                                        
  |  |                                            +-- Reginaia_ebenus **         
  |  |  +-----------------------------------------+                           
  +--+  |                                         +-- Reginaia_rotulata **       
     |  |                                                                     
     |  |                             +-------------- Parvaspina_collina **      
     |  |                             |                                       
     |  |                             |           +-- Eurynia_dilatata **        
     |  |  +--------------------------+  +--------+                           
     +--+  |                          |  |        +-- Hemistena_lata **          
        |  |                          |  |                                    
        |  |                          +--+        +-- Pleurobema_clava **        
        |  |                             |  +-----+                           
        |  |                             |  |     +-- Pleurobema_sintoxia **     
        |  |                             +--+                                 
        |  |                                |  +----- Pleuronaia_dolabelloides **
        +--+                                +--+                              
           |                                   |  +-- Fusconaia_flava **         
           |                                   +--+                           
           |                                      +-- Fusconaia_subrotunda **    
           |                                                                  
           |                                      +-- Amblema_plicata **         
           |  +-----------------------------------+                           
           |  |                                   +-- Amblema_elliottii **       
           |  |                                                               
           +--+  +----------------------------------- Toxolasma_lividum **       
              |  |                                                            
              |  |  +-------------------------------- Ellipsaria_lineolata **    
              +--+  |                                                         
                 |  |                          +----- Truncilla_cognata **       
                 |  |  +-----------------------+                              
                 |  |  |                       |  +-- Truncilla_macrodon **      
                 +--+  |                       +--+                           
                    |  |                          +-- Truncilla_donaciformis **  
                    |  |                                                      
                    |  |                    +-------- Potamilus_fragilis **      
                    |  |                    |                                 
                    +--+  +-----------------+     +-- Potamilus_ohiensis **      
                       |  |                 |  +--+                           
                       |  |                 |  |  +-- Potamilus_amphichaenus **  
                       |  |                 +--+                              
                       |  |                    |  +-- Potamilus_metnecktayi **   
                       |  |                    +--+                           
                       |  |                       +-- Potamilus_purpuratus **    
                       +--+                                                   
                          |  +----------------------- Leaunio_lienosus **        
                          |  |                                                
                          |  |     +----------------- Lampsilis_bracteata **     
                          |  |     |                                          
                          |  |  +--+              +-- Lampsilis_ornata **        
                          |  |  |  |  +-----------+                           
                          +--+  |  |  |           +-- Lampsilis_fasciola **      
                             |  |  +--+                                       
                             |  |     |  +----------- Lampsilis_ovata **         
                             |  |     +--+                                    
                             |  |        |  +-------- Lampsilis_cardium[3] **    
                             |  |        +--+                                 
                             +--+           |  +----- Lampsilis_cardium[2] **    
                                |           +--+                              
                                |              |  +-- Lampsilis_satura **        
                                |              +--+                           
                                |                 +-- Lampsilis_cardium[1] **    
                                |                                             
                                |     +-------------- Lampsilis_teres **         
                                |     |                                       
                                |     |           +-- Lampsilis_higginsii **     
                                +-----+  +--------+                           
                                      |  |        +-- Ortmanniana_ligamentina ** 
                                      |  |                                    
                                      +--+        +-- Lampsilis_radiata **       
                                         |  +-----+                           
                                         |  |     +-- Lampsilis_siliquoidea **   
                                         |  |                                 
                                         +--+     +-- Lampsilis_hydiana **       
                                            |  +--+                           
                                            |  |  +-- Lampsilis_bergmanni **     
                                            +--+                              
                                               |  +-- Lampsilis_virescens **     
                                               +--+                           
                                                  +-- Lampsilis_straminea **     

Dai et al., 2023 [Publ 4285], fig. 3-BI

iT tT bi riS oT roT roS iD
43 43 39 1 2 2 1 81.0
                          +-- Gibbosula_laosensis          
  +-----------------------+                                
  |                       +-- Margaritifera_margaritifera  
  |                                                        
  |                    +----- Indopseudodon_indawgyiensis **  
  |           +--------+                                   
  |           |        |  +-- Indopseudodon_bogani **         
  |           |        +--+                                
  |           |           +-- Indopseudodon_salwenianus **    
  |           |                                            
  |           |     +-------- Nyeinchanconcha_nyeinchani **   
  |           |  +--+                                      
  |           |  |  |  +----- Bineurus_loeiensis **           
  |  +--------+  |  +--+                                   
  |  |        |  |     |  +-- Bineurus_mouhotii **            
  |  |        |  |     +--+                                
--+  |        |  |        +-- Bineurus_exilis **              
  |  |        |  |                                         
  |  |        |  |        +-- Thaiconcha_callifera **         
  |  |        |  |  +-----+                                
  |  |        |  |  |     +-- Thaiconcha_munelliptica **      
  |  |        +--+--+                                      
  |  |           |  |  +----- Sundadontina_tanintharyiensis **
  |  |           |  +--+                                   
  |  |           |     |  +-- Sundadontina_brandti **         
  |  |           |     +--+                                
  |  |           |        +-- Sundadontina_taskaevi **        
  |  |           |                                         
  |  |           |        +-- Pilsbryoconcha_carinifera **    
  |  |           |  +-----+                                
  |  |           |  |     +-- Pilsbryoconcha_exilis **        
  |  |           +--+                                      
  +--+              |  +----- Pseudodon_lenyanensis **        
     |              +--+                                   
     |                 |  +-- Pseudodon_mekongi **            
     |                 +--+                                
     |                    +-- Pseudodon_vagulus **            
     |                                                     
     |                    +-- Hyriopsis_bialata **            
     |              +-----+                                
     |              |     +-- Ensidens_ingallsianus **        
     |  +-----------+                                      
     |  |           |  +----- Physunio_superbus **            
     |  |           +--+                                   
     |  |              |  +-- Lens_contradens **              
     |  |              +--+                                
     |  |                 +-- Lens_comptus **                 
     |  |                                                  
     |  |                 +-- Inversidens_brandtii **         
     |  |  +--------------+                                
     |  |  |              +-- Inversidens_rentianensis **     
     +--+  |                                               
        |  |           +----- Chamberlainia_somsakpanhai **   
        |  |     +-----+                                   
        |  |     |     |  +-- Sinohyriopsis_cumingii **       
        |  |     |     +--+                                
        |  +-----+        +-- Sinohyriopsis_schlegelii **     
        |  |     |                                         
        |  |     |  +-------- Potomida_littoralis **          
        |  |     +--+                                      
        |  |        |  +----- Pronodularia_japanensis **      
        +--+        +--+                                   
           |           |  +-- Lamprotula_leaii **             
           |           +--+                                
           |              +-- Lamprotula_caveata **           
           |                                               
           |        +-------- Sinosolenaia_carinata **        
           |  +-----+                                      
           |  |     |  +----- Sinosolenaia_recognita **       
           |  |     +--+                                   
           |  |        |  +-- Sinosolenaia_iridinea **        
           |  |        +--+                                
           |  |           +-- Sinosolenaia_oleivora **        
           +--+                                            
              |        +----- Gonidea_angulata **             
              |  +-----+                                   
              |  |     |  +-- Leguminaia_wheatleyi **         
              |  |     +--+                                
              |  |        +-- Microcondylaea_bonellii **      
              +--+                                         
                 |        +-- Koreosolenaia_sitgyensis **     
                 |  +-----+                                
                 |  |     +-- Parvasolenaia_rivularis **      
                 +--+                                      
                    |  +----- Postolata_guangxiensis **       
                    +--+                                   
                       |  +-- Ptychorhynchus_pfisteri **      
                       +--+                                
                          +-- Obovalis_omiensis **            

Dai et al., 2023 [Publ 4285], fig. 3-ML

iT tT bi riS oT roT roS iD
43 43 41 1 2 2 1 81.0
                          +-- Gibbosula_laosensis          
  +-----------------------+                                
  |                       +-- Margaritifera_margaritifera  
  |                                                        
  |                    +----- Indopseudodon_indawgyiensis **  
  |        +-----------+                                   
  |        |           |  +-- Indopseudodon_bogani **         
  |        |           +--+                                
  |        |              +-- Indopseudodon_salwenianus **    
  |        |                                               
  |        |        +-------- Nyeinchanconcha_nyeinchani **   
  |  +-----+  +-----+                                      
  |  |     |  |     |  +----- Bineurus_loeiensis **           
  |  |     |  |     +--+                                   
  |  |     |  |        |  +-- Bineurus_mouhotii **            
  |  |     |  |        +--+                                
--+  |     |  |           +-- Bineurus_exilis **              
  |  |     |  |                                            
  |  |     +--+           +-- Thaiconcha_callifera **         
  |  |        |     +-----+                                
  |  |        |     |     +-- Thaiconcha_munelliptica **      
  |  |        |  +--+                                      
  |  |        |  |  |  +----- Sundadontina_tanintharyiensis **
  |  |        |  |  +--+                                   
  |  |        |  |     |  +-- Sundadontina_brandti **         
  |  |        |  |     +--+                                
  |  |        +--+        +-- Sundadontina_taskaevi **        
  |  |           |                                         
  |  |           |        +-- Pilsbryoconcha_carinifera **    
  |  |           |  +-----+                                
  |  |           |  |     +-- Pilsbryoconcha_exilis **        
  +--+           +--+                                      
     |              |  +----- Pseudodon_lenyanensis **        
     |              +--+                                   
     |                 |  +-- Pseudodon_mekongi **            
     |                 +--+                                
     |                    +-- Pseudodon_vagulus **            
     |                                                     
     |                    +-- Hyriopsis_bialata **            
     |              +-----+                                
     |              |     +-- Ensidens_ingallsianus **        
     |  +-----------+                                      
     |  |           |  +----- Physunio_superbus **            
     |  |           +--+                                   
     |  |              |  +-- Lens_contradens **              
     |  |              +--+                                
     |  |                 +-- Lens_comptus **                 
     |  |                                                  
     |  |                 +-- Inversidens_brandtii **         
     |  |     +-----------+                                
     |  |     |           +-- Inversidens_rentianensis **     
     +--+     |                                            
        |  +--+        +----- Chamberlainia_somsakpanhai **   
        |  |  |  +-----+                                   
        |  |  |  |     |  +-- Sinohyriopsis_cumingii **       
        |  |  |  |     +--+                                
        |  |  +--+        +-- Sinohyriopsis_schlegelii **     
        |  |     |                                         
        |  |     |  +-------- Potomida_littoralis **          
        |  |     +--+                                      
        |  |        |  +----- Pronodularia_japanensis **      
        |  |        +--+                                   
        +--+           |  +-- Lamprotula_leaii **             
           |           +--+                                
           |              +-- Lamprotula_caveata **           
           |                                               
           |        +-------- Sinosolenaia_carinata **        
           |  +-----+                                      
           |  |     |  +----- Sinosolenaia_recognita **       
           |  |     +--+                                   
           |  |        |  +-- Sinosolenaia_iridinea **        
           |  |        +--+                                
           |  |           +-- Sinosolenaia_oleivora **        
           +--+                                            
              |        +----- Gonidea_angulata **             
              |  +-----+                                   
              |  |     |  +-- Leguminaia_wheatleyi **         
              |  |     +--+                                
              |  |        +-- Microcondylaea_bonellii **      
              +--+                                         
                 |        +-- Koreosolenaia_sitgyensis **     
                 |  +-----+                                
                 |  |     +-- Parvasolenaia_rivularis **      
                 +--+                                      
                    |  +----- Postolata_guangxiensis **       
                    +--+                                   
                       |  +-- Ptychorhynchus_pfisteri **      
                       +--+                                
                          +-- Obovalis_omiensis **            

Huang et al., 2019 [Publ 3843], fig. 1

iT tT bi riS oT roT roS iD
42 42 39 2 2 2 1 80.0
                                         +-- Margaritifera_dahurica     
  +--------------------------------------+                              
  |                                      +-- Gibbosula_rochechouartii   
  |                                                                     
  |                                   +----- Lamprotula_leaii **           
  |                                   |                                 
  |                       +-----------+  +-- Lamprotula_caveata[1] **      
  |                       |           |  |                              
  |                       |           +--+-- Lamprotula_cornuumlunae **    
  |                       |              |                              
--+  +--------------------+              +-- Lamprotula_caveata[2] **      
  |  |                    |                                             
  |  |                    |              +-- Parvasolenaia_rivularis **    
  |  |                    |  +-----------+                              
  |  |                    |  |           +-- Parvasolenaia_triangularis ** 
  |  |                    +--+                                          
  |  |                       |  +----------- Ptychorhynchus_pfisteri **    
  |  |                       +--+                                       
  |  |                          |  +-------- Sinosolenaia_carinata **      
  |  |                          +--+                                    
  +--+                             |  +----- Sinosolenaia_recognita **     
     |                             +--+                                 
     |                                |  +-- Sinosolenaia_oleivora **      
     |                                +--+                              
     |                                   +-- Sinosolenaia_iridinea **      
     |                                                                  
     |                                +----- Sinohyriopsis_cumingii[1] **  
     |  +-----------------------------+                                 
     |  |                             |  +-- Sinohyriopsis_cumingii[2] **  
     |  |                             +--+                              
     |  |                                +-- Sinohyriopsis_schlegelii **   
     +--+                                                               
        |  +-------------------------------- Anemina_arcaeformis **        
        |  |                                                            
        |  |  +----------------------------- Acuticosta_chinensis **       
        |  |  |                                                         
        +--+  |                          +-- Sinanodonta_angula **         
           |  |                 +--------+                              
           |  |                 |        +-- Sinanodonta_lucida **         
           |  |  +--------------+                                       
           |  |  |              |  +-------- Sinanodonta_jourdyi **        
           +--+  |              +--+                                    
              |  |                 |  +----- Sinanodonta_woodiana[2] **    
              |  |                 +--+                                 
              |  |                    |  +-- Sinanodonta_elliptica **      
              |  |                    +--+                              
              |  |                       +-- Sinanodonta_woodiana[1] **    
              |  |                                                      
              +--+              +----------- Aculamprotula_tortuosa **     
                 |  +-----------+                                       
                 |  |           |  +-------- Aculamprotula_scripta **      
                 |  |           +--+                                    
                 |  |              |  +----- Aculamprotula_coreana **      
                 |  |              +--+                                 
                 |  |                 |  +-- Aculamprotula_fibrosa **      
                 |  |                 +--+                              
                 |  |                    +-- Aculamprotula_tientsinensis **
                 +--+                                                   
                    |           +----------- Cristaria_plicata **          
                    |  +--------+                                       
                    |  |        |  +-------- Lanceolaria_gladiola **       
                    |  |        +--+                                    
                    |  |           |  +----- Lanceolaria_grayii **         
                    |  |           +--+                                 
                    |  |              |  +-- Lanceolaria_lanceolata **     
                    +--+              +--+                              
                       |                 +-- Lanceolaria_triformis **      
                       |                                                
                       |  +----------------- Lepidodesma_languilati **     
                       |  |                                             
                       |  |              +-- Schistodesmus_lampreyanus **  
                       +--+  +-----------+                              
                          |  |           +-- Schistodesmus_spinosus **     
                          |  |                                          
                          +--+        +----- Nodularia_douglasiae[1] **    
                             |  +-----+                                 
                             |  |     |  +-- Nodularia_douglasiae[2] **    
                             |  |     +--+                              
                             +--+        +-- Nodularia_nuxpersicae **      
                                |                                       
                                |  +-------- Cuneopsis_rufescens **        
                                +--+                                    
                                   |  +----- Tchangsinaia_piscicula **     
                                   +--+                                 
                                      |  +-- Cuneopsis_heudei **           
                                      +--+                              
                                         +-- Cuneopsis_celtiformis **      

Froufe et al., 2019 [Publ 3936], fig. 3

iT tT bi riS oT roT roS iD
42 42 40 3 3 3 1 78.0
                             +----- Pseudunio_marocanus        
  +--------------------------+                                 
  |                          |  +-- Margaritifera_dahurica     
  |                          +--+                              
  |                             +-- Margaritifera_falcata      
  |                                                            
  |                    +----------- Quadrula_quadrula **          
  |                    |                                       
  |  +-----------------+  +-------- Toxolasma_parvum **           
  |  |                 |  |                                    
  |  |                 +--+     +-- Lampsilis_ornata **           
  |  |                    |  +--+                              
--+  |                    |  |  +-- Venustaconcha_ellipsiformis **
  |  |                    +--+                                 
  |  |                       |  +-- Potamilus_leptodon **         
  |  |                       +--+                              
  |  |                          +-- Potamilus_alatus **           
  |  |                                                         
  |  |                       +----- Chamberlainia_somsakpanhai ** 
  |  |           +-----------+                                 
  |  |           |           |  +-- Sinohyriopsis_cumingii **     
  |  |           |           +--+                              
  |  |           |              +-- Sinohyriopsis_schlegelii **   
  +--+           |                                             
     |  +--------+        +-------- Potomida_littoralis **        
     |  |        |  +-----+                                    
     |  |        |  |     |  +----- Pronodularia_japanensis **    
     |  |        |  |     +--+                                 
     |  |        |  |        |  +-- Lamprotula_leaii **           
     |  |        |  |        +--+                              
     |  |        +--+           +-- Lamprotula_caveata **         
     |  |           |                                          
     |  |           |           +-- Pilsbryoconcha_exilis **      
     |  |           |  +--------+                              
     |  |           |  |        +-- Pseudodon_vondembuschianus ** 
     |  |           +--+                                       
     |  |              |  +-------- Ptychorhynchus_pfisteri **    
     |  |              +--+                                    
     +--+                 |  +----- Microcondylaea_bonellii **    
        |                 +--+                                 
        |                    |  +-- Sinosolenaia_carinata **      
        |                    +--+                              
        |                       +-- Sinosolenaia_oleivora **      
        |                                                      
        |  +----------------------- Lepidodesma_languilati **     
        |  |                                                   
        |  |                 +----- Aculamprotula_tortuosa **     
        |  |        +--------+                                 
        |  |        |        |  +-- Aculamprotula_tientsinensis **
        |  |        |        +--+                              
        |  |        |           +-- Aculamprotula_coreana **      
        |  |  +-----+                                          
        +--+  |     |        +----- Nodularia_douglasiae **       
           |  |     |  +-----+                                 
           |  |     |  |     |  +-- Tchangsinaia_piscicula **     
           |  |     |  |     +--+                              
           |  |     +--+        +-- Schistodesmus_lampreyanus **  
           |  |        |                                       
           |  |        |  +-------- Unio_tumidus **               
           |  |        +--+                                    
           |  |           |  +----- Unio_crassus **               
           +--+           +--+                                 
              |              |  +-- Unio_pictorum **              
              |              +--+                              
              |                 +-- Unio_delphinus **             
              |                                                
              |                 +-- Lanceolaria_lanceolata **     
              |  +--------------+                              
              |  |              +-- Lanceolaria_grayii **         
              |  |                                             
              |  |        +-------- Cristaria_plicata **          
              +--+        |                                    
                 |  +-----+     +-- Anemina_arcaeformis **        
                 |  |     |  +--+                              
                 |  |     |  |  +-- Anemina_euscaphys **          
                 |  |     +--+                                 
                 +--+        |  +-- Sinanodonta_lucida **         
                    |        +--+                              
                    |           +-- Sinanodonta_woodiana **       
                    |                                          
                    |  +----------- Anodonta_anatina **           
                    +--+                                       
                       |  +-------- Lasmigona_complanata **       
                       +--+                                    
                          |  +----- Pyganodon_grandis **          
                          +--+                                 
                             |  +-- Utterbackia_peninsularis **   
                             +--+                              
                                +-- Utterbackia_imbecillis **     

Froufe et al., 2019 [Publ 3936], fig. 6

iT tT bi riS oT roT roS iD
42 42 40 3 3 3 1 78.0
                             +----- Pseudunio_marocanus        
  +--------------------------+                                 
  |                          |  +-- Margaritifera_dahurica     
  |                          +--+                              
  |                             +-- Margaritifera_falcata      
  |                                                            
  |                    +----------- Quadrula_quadrula **          
  |                    |                                       
  |  +-----------------+  +-------- Toxolasma_parvum **           
  |  |                 |  |                                    
  |  |                 +--+     +-- Potamilus_leptodon **         
  |  |                    |  +--+                              
--+  |                    |  |  +-- Potamilus_alatus **           
  |  |                    +--+                                 
  |  |                       |  +-- Lampsilis_ornata **           
  |  |                       +--+                              
  |  |                          +-- Venustaconcha_ellipsiformis **
  |  |                                                         
  |  |                       +----- Chamberlainia_somsakpanhai ** 
  |  |           +-----------+                                 
  |  |           |           |  +-- Sinohyriopsis_cumingii **     
  |  |           |           +--+                              
  |  |           |              +-- Sinohyriopsis_schlegelii **   
  +--+           |                                             
     |  +--------+        +-------- Potomida_littoralis **        
     |  |        |  +-----+                                    
     |  |        |  |     |  +----- Pronodularia_japanensis **    
     |  |        |  |     +--+                                 
     |  |        |  |        |  +-- Lamprotula_leaii **           
     |  |        |  |        +--+                              
     |  |        +--+           +-- Lamprotula_caveata **         
     |  |           |                                          
     |  |           |           +-- Pilsbryoconcha_exilis **      
     |  |           |  +--------+                              
     |  |           |  |        +-- Pseudodon_vondembuschianus ** 
     |  |           +--+                                       
     |  |              |  +-------- Ptychorhynchus_pfisteri **    
     |  |              +--+                                    
     +--+                 |  +----- Microcondylaea_bonellii **    
        |                 +--+                                 
        |                    |  +-- Sinosolenaia_carinata **      
        |                    +--+                              
        |                       +-- Sinosolenaia_oleivora **      
        |                                                      
        |  +----------------------- Lepidodesma_languilati **     
        |  |                                                   
        |  |                 +----- Aculamprotula_tortuosa **     
        |  |        +--------+                                 
        |  |        |        |  +-- Aculamprotula_tientsinensis **
        |  |        |        +--+                              
        |  |        |           +-- Aculamprotula_coreana **      
        |  |  +-----+                                          
        +--+  |     |        +----- Nodularia_douglasiae **       
           |  |     |  +-----+                                 
           |  |     |  |     |  +-- Tchangsinaia_piscicula **     
           |  |     |  |     +--+                              
           |  |     +--+        +-- Schistodesmus_lampreyanus **  
           |  |        |                                       
           |  |        |  +-------- Unio_tumidus **               
           |  |        +--+                                    
           |  |           |  +----- Unio_crassus **               
           +--+           +--+                                 
              |              |  +-- Unio_pictorum **              
              |              +--+                              
              |                 +-- Unio_delphinus **             
              |                                                
              |                 +-- Lanceolaria_lanceolata **     
              |  +--------------+                              
              |  |              +-- Lanceolaria_grayii **         
              |  |                                             
              |  |     +----------- Anodonta_anatina **           
              +--+  +--+                                       
                 |  |  |  +-------- Lasmigona_complanata **       
                 |  |  +--+                                    
                 |  |     |  +----- Pyganodon_grandis **          
                 |  |     +--+                                 
                 +--+        |  +-- Utterbackia_peninsularis **   
                    |        +--+                              
                    |           +-- Utterbackia_imbecillis **     
                    |                                          
                    |     +-------- Cristaria_plicata **          
                    |     |                                    
                    +-----+     +-- Sinanodonta_lucida **         
                          |  +--+                              
                          |  |  +-- Sinanodonta_woodiana **       
                          +--+                                 
                             |  +-- Anemina_arcaeformis **        
                             +--+                              
                                +-- Anemina_euscaphys **          

Smith et al., 2019 [Publ 3951], fig. 2

iT tT bi riS oT roT roS iD
42 42 40 1 0 0 0 77.0
  +----------------------------------------------- Quadrula_quadrula **          
  |                                                                           
  |                                            +-- Amblema_plicata **            
--+  +-----------------------------------------+                              
  |  |                                         +-- Reginaia_ebenus **            
  |  |                                                                        
  +--+  +----------------------------------------- Plectomerus_dombeyanus **     
     |  |                                                                     
     |  |  +-------------------------------------- Toxolasma_lividum **          
     +--+  |                                                                  
        |  |                                +----- Popenaias_popeii **           
        |  |  +-----------------------------+                                 
        +--+  |                             |  +-- Glebula_rotundata **          
           |  |                             +--+                              
           |  |                                +-- Cyrtonaias_tampicoensis **    
           |  |                                                               
           |  |  +-------------------------------- Obliquaria_reflexa **         
           +--+  |                                                            
              |  |                             +-- Medionidus_conradicus **      
              |  |                          +--+                              
              |  |                          |  +-- Lemiox_rimosus **             
              |  |  +-----------------------+                                 
              |  |  |                       |  +-- Cyprogenia_stegaria **        
              |  |  |                       +--+                              
              +--+  |                          +-- Ptychobranchus_fasciolaris ** 
                 |  |                                                         
                 |  |                          +-- Hamiota_subangulata **        
                 |  |     +--------------------+                              
                 |  |     |                    +-- Villosa_villosa **            
                 |  |     |                                                   
                 |  |  +--+  +-------------------- Ligumia_recta **              
                 |  |  |  |  |                                                
                 +--+  |  |  |              +----- Venustaconcha_ellipsiformis **
                    |  |  +--+  +-----------+                                 
                    |  |     |  |           |  +-- Obovaria_subrotunda **        
                    |  |     |  |           +--+                              
                    |  |     +--+              +-- Epioblasma_rangiana **        
                    |  |        |                                             
                    |  |        |  +-------------- Ortmanniana_pectorosa **      
                    |  |        +--+                                          
                    |  |           |  +----------- Ortmanniana_ligamentina **    
                    |  |           +--+                                       
                    |  |              |  +-------- Lampsilis_ovata **            
                    |  |              +--+                                    
                    +--+                 |  +----- Lampsilis_dolabraeformis **   
                       |                 +--+                                 
                       |                    |  +-- Lampsilis_satura **           
                       |                    +--+                              
                       |                       +-- Lampsilis_cardium **          
                       |                                                      
                       |           +-------------- Atlanticoncha_ochracea **     
                       |     +-----+                                          
                       |     |     |  +----------- Ellipsaria_lineolata **       
                       |     |     +--+                                       
                       |     |        |  +-------- Truncilla_truncata **         
                       |     |        +--+                                    
                       |     |           |  +----- Truncilla_cognata **          
                       |     |           +--+                                 
                       +-----+              |  +-- Truncilla_macrodon **         
                             |              +--+                              
                             |                 +-- Truncilla_donaciformis **     
                             |                                                
                             |  +----------------- Potamilus_capax **            
                             |  |                                             
                             |  |           +----- Potamilus_inflatus **         
                             +--+  +--------+                                 
                                |  |        |  +-- Potamilus_fragilis **         
                                |  |        +--+                              
                                |  |           +-- Potamilus_leptodon **         
                                +--+                                          
                                   |        +----- Potamilus_ohiensis **         
                                   |  +-----+                                 
                                   |  |     |  +-- Potamilus_amphichaenus **     
                                   |  |     +--+                              
                                   +--+        +-- Potamilus_streckersoni **     
                                      |                                       
                                      |  +-------- Potamilus_metnecktayi **      
                                      +--+                                    
                                         |  +----- Potamilus_purpuratus[1] **    
                                         +--+                                 
                                            |  +-- Potamilus_purpuratus[2] **    
                                            +--+                              
                                               +-- Potamilus_alatus **           

Wu et al., 2020 [Publ 4121], fig. 3

iT tT bi riS oT roT roS iD
42 42 40 3 8 4 1 81.0
                          +-------- Gibbosula_rochechouartii   
  +-----------------------+                                    
  |                       |  +----- Cumberlandia_monodonta     
  |                       +--+                                 
  |                          |  +-- Margaritifera_falcata      
  |                          +--+                              
  |                             +-- Margaritifera_dahurica     
  |                                                            
  |                    +----------- Quadrula_quadrula **          
  |                    |                                       
  |  +-----------------+  +-------- Toxolasma_parvum **           
--+  |                 |  |                                    
  |  |                 +--+     +-- Venustaconcha_ellipsiformis **
  |  |                    |  +--+                              
  |  |                    |  |  +-- Lampsilis_ornata **           
  |  |                    +--+                                 
  |  |                       |  +-- Potamilus_alatus **           
  |  |                       +--+                              
  |  |                          +-- Potamilus_leptodon **         
  |  |                                                         
  |  |                          +-- Sinohyriopsis_schlegelii **   
  |  |                 +--------+                              
  +--+                 |        +-- Sinohyriopsis_cumingii **     
     |                 |                                       
     |  +--------------+     +----- Potomida_littoralis **        
     |  |              |  +--+                                 
     |  |              |  |  |  +-- Pronodularia_japanensis **    
     |  |              |  |  +--+                              
     |  |              +--+     +-- Lamprotula_leaii **           
     |  |                 |                                    
     |  |                 |  +----- Ptychorhynchus_pfisteri **    
     |  |                 +--+                                 
     |  |                    |  +-- Sinosolenaia_oleivora **      
     |  |                    +--+                              
     |  |                       +-- Sinosolenaia_carinata **      
     +--+                                                      
        |  +----------------------- Lepidodesma_languilati **     
        |  |                                                   
        |  |                 +----- Acuticosta_chinensis **       
        |  |     +-----------+                                 
        |  |     |           |  +-- Lanceolaria_lanceolata **     
        |  |     |           +--+                              
        |  |     |              +-- Lanceolaria_grayii **         
        |  |     |                                             
        |  |  +--+     +----------- Anodonta_anatina **           
        |  |  |  |  +--+                                       
        |  |  |  |  |  |  +-------- Platynaias_compressa **       
        +--+  |  |  |  +--+                                    
           |  |  |  |     |  +----- Pyganodon_grandis **          
           |  |  |  |     +--+                                 
           |  |  +--+        |  +-- Utterbackia_peninsularis **   
           |  |     |        +--+                              
           |  |     |           +-- Utterbackia_imbecillis **     
           |  |     |                                          
           |  |     |     +-------- Cristaria_plicata **          
           |  |     |     |                                    
           |  |     +-----+     +-- Sinanodonta_woodiana **       
           |  |           |  +--+                              
           +--+           |  |  +-- Sinanodonta_lauta **          
              |           +--+                                 
              |              |  +-- Anemina_arcaeformis **        
              |              +--+                              
              |                 +-- Anemina_euscaphys **          
              |                                                
              |           +-------- Aculamprotula_tortuosa **     
              |           |                                    
              |  +--------+     +-- Aculamprotula_coreana **      
              |  |        |  +--+                              
              |  |        |  |  +-- Aculamprotula_tientsinensis **
              |  |        +--+                                 
              |  |           |  +-- Aculamprotula_polysticta **   
              |  |           +--+                              
              +--+              +-- Aculamprotula_scripta **      
                 |                                             
                 |        +-------- Unio_tumidus **               
                 |  +-----+                                    
                 |  |     |  +----- Unio_crassus **               
                 |  |     +--+                                 
                 |  |        |  +-- Unio_delphinus **             
                 +--+        +--+                              
                    |           +-- Unio_pictorum **              
                    |                                          
                    |  +----------- Pseudocuneopsis_capitata **   
                    +--+                                       
                       |  +-------- Nodularia_douglasiae **       
                       +--+                                    
                          |  +----- Tchangsinaia_piscicula **     
                          +--+                                 
                             |  +-- Cuneopsis_heudei **           
                             +--+                              
                                +-- Schistodesmus_lampreyanus **  

Bolotov et al., 2023 [Publ 4312], fig. S2

iT tT bi riS oT roT roS iD
42 42 39 1 0 0 0 82.0
                          +----------- Indopseudodon_kayinensis **       
  +-----------------------+                                           
  |                       |  +-------- Indopseudodon_indawgyiensis **    
  |                       +--+                                        
  |                          |  +----- Indopseudodon_bogani **           
  |                          +--+                                     
  |                             |  +-- Indopseudodon_salwenianus **      
  |                             +--+                                  
  |                                +-- Indopseudodon_crebristriatus **   
  |                                                                   
--+  +-------------------------------- Songkhlanaia_tamodienica **       
  |  |                                                                
  |  |              +----------------- Nyeinchanconcha_nyeinchani **     
  |  |              |                                                 
  |  |  +-----------+  +-------------- Sundadontina_tumida **            
  |  |  |           |  |                                              
  |  |  |           |  |     +-------- Sundadontina **                   
  |  |  |           +--+  +--+                                        
  |  |  |              |  |  |  +----- Sundadontina_tanintharyiensis **  
  |  |  |              |  |  +--+                                     
  +--+  |              |  |     |  +-- Sundadontina_plugpomenica **      
     |  |              +--+     +--+                                  
     |  |                 |        +-- Sundadontina_cumingii **          
     |  |                 |                                           
     |  |                 |  +-------- Sundadontina_sulcata **           
     |  |                 +--+                                        
     |  |                    |  +----- Sundadontina_harmandi **          
     |  |                    +--+                                     
     |  |                       |  +-- Sundadontina_taskaevi **          
     |  |                       +--+                                  
     |  |                          +-- Sundadontina_brandti **           
     +--+                                                             
        |                          +-- Namkongnaia_inkhavilayi **        
        |           +--------------+                                  
        |           |              +-- Namkongnaia_lemeslei **           
        |           |                                                 
        |  +--------+              +-- Thaiconcha_callifera **           
        |  |        |              |                                  
        |  |        |  +-----------+-- Thaiconcha_thaiensis **           
        |  |        |  |           |                                  
        |  |        +--+           +-- Thaiconcha_munelliptica **        
        |  |           |                                              
        |  |           |  +----------- Bineurus_loeiensis **             
        |  |           +--+                                           
        |  |              |  +-------- Bineurus_panhai **                
        |  |              +--+                                        
        |  |                 |  +----- Bineurus_anodontinum **           
        |  |                 +--+                                     
        +--+                    |  +-- Bineurus_exilis **                
           |                    +--+                                  
           |                       +-- Bineurus_mouhotii **              
           |                                                          
           |              +----------- Pseudodon_vagulus **              
           |              |                                           
           |  +-----------+        +-- Pseudodon_lenyanensis **          
           |  |           |  +-----+                                  
           |  |           |  |     +-- Pseudodon_inoscularis **          
           |  |           +--+                                        
           |  |              |  +----- Pseudodon_cambodjensis **         
           |  |              +--+                                     
           +--+                 |  +-- Pseudodon_vondembuschianus **     
              |                 +--+                                  
              |                    +-- Pseudodon_mekongi **              
              |                                                       
              |  +-------------------- Pilsbryoconcha_carinifera **      
              |  |                                                    
              +--+                 +-- Pilsbryoconcha_linguaeformis[2] **
                 |  +--------------+                                  
                 |  |              +-- Pilsbryoconcha **                 
                 +--+                                                 
                    |  +-------------- Pilsbryoconcha_linguaeformis[1] **
                    |  |                                              
                    +--+           +-- Pilsbryoconcha_schomburgki **     
                       |  +--------+                                  
                       |  |        +-- Pilsbryoconcha_exilis **          
                       +--+                                           
                          |  +-------- Pilsbryoconcha_hoikaab **         
                          +--+                                        
                             |  +----- Pilsbryoconcha_mekongiana **      
                             +--+                                     
                                |  +-- Pilsbryoconcha_acuta **           
                                +--+                                  
                                   +-- Pilsbryoconcha_kittitati **       

Wu et al., 2019 [Publ 3864], fig. 3

iT tT bi riS oT roT roS iD
41 41 39 3 6 4 1 80.0
                    +----------- Quadrula_quadrula **          
                    |                                       
  +-----------------+  +-------- Toxolasma_parvum **           
  |                 |  |                                    
  |                 +--+     +-- Lampsilis_ornata **           
  |                    |  +--+                              
  |                    |  |  +-- Venustaconcha_ellipsiformis **
  |                    +--+                                 
  |                       |  +-- Potamilus_leptodon **         
  |                       +--+                              
  |                          +-- Potamilus_alatus **           
  |                                                         
  |                          +-- Sinohyriopsis_cumingii **     
  |                 +--------+                              
--+                 |        +-- Sinohyriopsis_schlegelii **   
  |                 |                                       
  |  +--------------+     +----- Potomida_littoralis **        
  |  |              |  +--+                                 
  |  |              |  |  |  +-- Lamprotula_leaii **           
  |  |              |  |  +--+                              
  |  |              +--+     +-- Pronodularia_japanensis **    
  |  |                 |                                    
  |  |                 |  +----- Ptychorhynchus_pfisteri **    
  |  |                 +--+                                 
  |  |                    |  +-- Sinosolenaia_oleivora **      
  |  |                    +--+                              
  |  |                       +-- Sinosolenaia_carinata **      
  +--+                                                      
     |  +----------------------- Lepidodesma_languilati **     
     |  |                                                   
     |  |              +-------- Aculamprotula_tortuosa **     
     |  |     +--------+                                    
     |  |     |        |  +----- Aculamprotula_scripta **      
     |  |     |        +--+                                 
     |  |     |           |  +-- Aculamprotula_tientsinensis **
     |  |     |           +--+                              
     |  |     |              +-- Aculamprotula_coreana **      
     |  |  +--+                                             
     +--+  |  |        +-------- Unio_tumidus **               
        |  |  |  +-----+                                    
        |  |  |  |     |  +----- Unio_crassus **               
        |  |  |  |     +--+                                 
        |  |  |  |        |  +-- Unio_delphinus **             
        |  |  +--+        +--+                              
        |  |     |           +-- Unio_pictorum **              
        |  |     |                                          
        |  |     |  +----------- Pseudocuneopsis_capitata **   
        |  |     +--+                                       
        |  |        |  +-------- Nodularia_douglasiae **       
        +--+        +--+                                    
           |           |  +----- Tchangsinaia_piscicula **     
           |           +--+                                 
           |              |  +-- Cuneopsis_heudei **           
           |              +--+                              
           |                 +-- Schistodesmus_lampreyanus **  
           |                                                
           |              +----- Acuticosta_chinensis **       
           |  +-----------+                                 
           |  |           |  +-- Lanceolaria_grayii **         
           |  |           +--+                              
           |  |              +-- Lanceolaria_lanceolata **     
           |  |                                             
           +--+     +----------- Anodonta_anatina **           
              |  +--+                                       
              |  |  |  +-------- Platynaias_compressa **       
              |  |  +--+                                    
              |  |     |  +----- Pyganodon_grandis **          
              |  |     +--+                                 
              +--+        |  +-- Utterbackia_peninsularis **   
                 |        +--+                              
                 |           +-- Utterbackia_imbecillis **     
                 |                                          
                 |     +-------- Cristaria_plicata **          
                 |     |                                    
                 +-----+     +-- Anemina_arcaeformis **        
                       |  +--+                              
                       |  |  +-- Anemina_euscaphys **          
                       +--+                                 
                          |  +-- Sinanodonta_lucida **         
                          +--+                              
                             +-- Sinanodonta_woodiana **       

Burlakova et al., 2012 [Publ 3486], fig. 2

iT tT bi riS oT roT roS iD
40 40 30 1 0 0 0 77.0
  +----------------------------------- Parvaspina_collina **      
  |                                                            
  |  +-------------------------------- Pleurobema_plenum **       
  |  |                                                         
--+  |                             +-- Plethobasus_cyphyus **     
  |  |                       +-----+                           
  |  |                       |     +-- Hemistena_lata **          
  |  |  +--------------------+                                 
  +--+  |                    |  +----- Eurynia_dilatata **        
     |  |                    +--+                              
     |  |                       |  +-- Elliptio_arca **           
     |  |                       +--+                           
     |  |                          +-- Elliptio_crassidens **     
     |  |                                                      
     |  |  +-------------------------- Pleurobema_cordatum[2] **  
     +--+  |                                                   
        |  |           +-------------- Fusconaia_subrotunda **    
        |  |           |                                       
        |  |           |           +-- Fusconaia_cor **           
        |  |           |  +--------+                           
        |  |  +--------+  |        +-- Fusconaia_cuneolus **      
        |  |  |        |  |                                    
        |  |  |        |  |  +-------- Fusconaia_flava[2] **      
        |  |  |        |  |  |                                 
        +--+  |        +--+  +-------- Fusconaia_ozarkensis **    
           |  |           |  |                                 
           |  |           |  +-------- Fusconaia_masoni **        
           |  |           |  |                                 
           |  |           |  |  +----- Fusconaia_askewi **        
           |  |           +--+--+                              
           |  |              |  |  +-- Fusconaia_burkei **        
           |  |              |  +--+                           
           |  |              |     +-- Fusconaia_escambia **      
           |  |              |                                 
           +--+              |     +-- Fusconaia_flava[1] **      
              |              |     |                           
              |              +-----+-- Fusconaia_cerina[1] **     
              |                    |                           
              |                    +-- Fusconaia_cerina[2] **     
              |                                                
              |              +-------- Pleurobema_stabile **      
              |              |                                 
              |  +-----------+  +----- Pleurobema_riddellii **    
              |  |           |  |                              
              |  |           +--+  +-- Pleurobema_cordatum[1] **  
              |  |              |  |                           
              |  |              |  +-- Pleurobema_rubrum **       
              |  |              +--+                           
              |  |                 +-- Pleurobema_sintoxia[1] **  
              +--+                 |                           
                 |                 +-- Pleurobema_sintoxia[2] **  
                 |                                             
                 |              +----- Pleuronaia_barnesiana **   
                 |  +-----------+                              
                 |  |           |  +-- Pleuronaia_dolabelloides **
                 |  |           +--+                           
                 |  |              +-- Pleuronaia_gibber **       
                 +--+                                          
                    |              +-- Pleurobema_georgianum **   
                    |  +-----------+                           
                    |  |           +-- Pleurobema_pyriforme **    
                    |  |                                       
                    +--+           +-- Pleurobema_decisum **      
                       |  +--------+                           
                       |  |        +-- Pleurobema_hanleyianum **  
                       |  |                                    
                       +--+        +-- Pleurobema_clava **        
                          |  +-----+                           
                          |  |     +-- Pleurobema_oviforme **     
                          |  |                                 
                          +--+  +----- Pleurobema_rubellum **     
                             |  |                              
                             |  +----- Pleurobema_beadleianum **  
                             +--+                              
                                +----- Pleurobema_strodeanum **   
                                |                              
                                |  +-- Pleurobema_perovatum **    
                                +--+                           
                                   +-- Pleurobema_taitianum **    

Bolotov et al., 2023 [Publ 4312], fig. 2

iT tT bi riS oT roT roS iD
40 40 38 1 0 0 0 81.0
                    +----------- Indopseudodon_kayinensis **     
  +-----------------+                                         
  |                 |  +-------- Indopseudodon_indawgyiensis **  
  |                 +--+                                      
  |                    |  +----- Indopseudodon_bogani **         
  |                    +--+                                   
  |                       |  +-- Indopseudodon_salwenianus **    
  |                       +--+                                
  |                          +-- Indopseudodon_crebristriatus ** 
  |                                                           
  |                          +-- Pilsbryoconcha_exilis **        
  |              +-----------+                                
--+              |           +-- Pilsbryoconcha_schomburgki **   
  |              |                                            
  |  +-----------+           +-- Pilsbryoconcha_carinifera **    
  |  |           |  +--------+                                
  |  |           |  |        +-- Pilsbryoconcha_linguaeformis ** 
  |  |           +--+                                         
  |  |              |  +-------- Pilsbryoconcha_acuta **         
  |  |              +--+                                      
  |  |                 |  +----- Pilsbryoconcha_kittitati **     
  |  |                 +--+                                   
  |  |                    |  +-- Pilsbryoconcha_mekongiana **    
  |  |                    +--+                                
  +--+                       +-- Pilsbryoconcha_hoikaab **       
     |                                                        
     |  +----------------------- Songkhlanaia_tamodienica **     
     |  |                                                     
     |  |                    +-- Namkongnaia_inkhavilayi **      
     |  |           +--------+                                
     |  |           |        +-- Namkongnaia_lemeslei **         
     |  |           |                                         
     |  |           |     +----- Pseudodon_vondembuschianus[2] **
     |  |  +--------+  +--+                                   
     |  |  |        |  |  |  +-- Pseudodon_vondembuschianus[1] **
     +--+  |        |  |  +--+                                
        |  |        |  |     +-- Pseudodon_mekongi **            
        |  |        +--+                                      
        |  |           |     +-- Pseudodon_vagulus **            
        |  |           |  +--+                                
        |  |           |  |  +-- Pseudodon_cambodjensis **       
        |  |           +--+                                   
        |  |              |  +-- Pseudodon_lenyanensis **        
        |  |              +--+                                
        |  |                 +-- Pseudodon_inoscularis **        
        +--+                                                  
           |     +-------------- Nyeinchanconcha_nyeinchani **   
           |     |                                            
           |     |        +----- Thaiconcha_munelliptica **      
           |  +--+  +-----+                                   
           |  |  |  |     |  +-- Thaiconcha_callifera **         
           |  |  |  |     +--+                                
           |  |  +--+        +-- Thaiconcha_thaiensis **         
           |  |     |                                         
           |  |     |  +-------- Bineurus_loeiensis **           
           |  |     +--+                                      
           |  |        |  +----- Bineurus_mouhotii **            
           |  |        +--+                                   
           +--+           |  +-- Bineurus_anodontinum **         
              |           +--+                                
              |              +-- Bineurus_exilis **              
              |                                               
              |        +-------- Sundadontina_sulcata **         
              |  +-----+                                      
              |  |     |  +----- Sundadontina_harmandi **        
              |  |     +--+                                   
              |  |        |  +-- Sundadontina_brandti **         
              +--+        +--+                                
                 |           +-- Sundadontina_taskaevi **        
                 |                                            
                 |  +----------- Sundadontina_tumida **          
                 +--+                                         
                    |  +-------- Sundadontina **                 
                    +--+                                      
                       |  +----- Sundadontina_tanintharyiensis **
                       +--+                                   
                          |  +-- Sundadontina_cumingii **        
                          +--+                                
                             +-- Sundadontina_plugpomenica **    

Pfeiffer et al., 2016 [Publ 3660], fig. 2

iT tT bi riS oT roT roS iD
39 39 34 2 0 0 0 73.0
  +-------------------------------------- Utterbackiana_suborbiculata **
  |                                                                  
  |                                   +-- Amblema_plicata **            
  |                                +--+                              
  |                                |  +-- Amblema_neislerii **          
  |  +-----------------------------+                                 
--+  |                             |  +-- Reginaia_rotulata **          
  |  |                             +--+                              
  |  |                                +-- Reginaia_ebenus **            
  |  |                                                               
  |  |                                +-- Cyrtonaias_tampicoensis **    
  |  |                          +-----+                              
  |  |                          |     +-- Glebula_rotundata **          
  +--+                          |                                    
     |  +-----------------------+     +-- Lampsilis_teres **            
     |  |                       |  +--+                              
     |  |                       |  |  +-- Leaunio_lienosus **           
     |  |                       +--+                                 
     |  |                          |  +-- Truncilla_macrodon **         
     |  |                          +--+                              
     |  |                             +-- Potamilus_fragilis **         
     +--+                                                            
        |  +----------------------------- Elliptoideus_sloatianus **    
        |  |                                                         
        |  |              +-------------- Uniomerus_declivis **         
        |  |  +-----------+                                          
        |  |  |           |  +----------- Quadrula_quadrula **          
        |  |  |           +--+                                       
        +--+  |              |  +-------- Quadrula_verrucosa **         
           |  |              +--+                                    
           |  |                 |  +----- Pustulosa_pustulosa **        
           |  |                 +--+                                 
           |  |                    |  +-- Pustulosa_petrina **          
           |  |                    +--+                              
           +--+                       +-- Cyclonaias_tuberculata **     
              |                                                      
              |                       +-- Elliptio_fumata **            
              |  +--------------------+                              
              |  |                    +-- Elliptio_crassidens **        
              |  |                                                   
              |  |  +-------------------- Parvaspina_collina **         
              |  |  |                                                
              |  |  +-------------------- Plethobasus_cyphyus **        
              +--+  |                                                
                 |  |                 +-- Hemistena_lata **             
                 |  +-----------------+                              
                 |  |                 +-- Eurynia_dilatata **           
                 |  |                                                
                 |  |              +----- Pleurobema_riddellii **       
                 +--+           +--+                                 
                    |           |  |  +-- Pleurobema_sintoxia **        
                    |           |  +--+                              
                    |  +--------+     +-- Pleurobema_cordatum **        
                    |  |        |                                    
                    |  |        |  +----- Pleuronaia_dolabelloides **   
                    |  |        |  |                                 
                    |  |        +--+  +-- Pleurobema_pyriforme **       
                    |  |           |  |                              
                    +--+           +--+-- Pleurobema_strodeanum **      
                       |              |                              
                       |              +-- Pleurobema_clava **           
                       |                                             
                       |  +-------------- Fusconaia_askewi **           
                       |  |                                          
                       |  |           +-- Fusconaia_mitchelli **        
                       +--+  +--------+                              
                          |  |        +-- Fusconaia_iheringi **         
                          |  |                                       
                          +--+        +-- Fusconaia_escambia **         
                             |  +-----+                              
                             |  |     +-- Fusconaia_burkei **           
                             +--+                                    
                                |  +----- Fusconaia_flava[2] **         
                                +--+                                 
                                   |  +-- Fusconaia_cerina **           
                                   +--+                              
                                      +-- Fusconaia_flava[1] **         

Pfeiffer et al., 2019 [Publ 3866], fig. 1

iT tT bi riS oT roT roS iD
39 39 34 2 0 0 0 61.0
  +-------------------------------------- Gonidea_angulata **           
  |                                                                  
  |                          +----------- Megalonaias_nervosa **        
  |  +-----------------------+                                       
  |  |                       |  +-------- Uniomerus_declivis **         
  |  |                       +--+                                    
--+  |                          |  +----- Quadrula_verrucosa **         
  |  |                          +--+                                 
  |  |                             |  +-- Cyclonaias_tuberculata **     
  |  |                             +--+                              
  |  |                                +-- Quadrula_quadrula **          
  |  |                                                               
  |  |                    +-------------- Elliptoideus_sloatianus **    
  +--+  +-----------------+                                          
     |  |                 |  +----------- Elliptio_fumata **            
     |  |                 +--+                                       
     |  |                    |  +-------- Pleuronaia_barnesiana **      
     |  |                    +--+                                    
     |  |                       |  +----- Pleurobema_riddellii **       
     |  |                       +--+                                 
     |  |                          |  +-- Fusconaia_flava **            
     |  |                          +--+                              
     |  |                             +-- Pleurobema_strodeanum **      
     |  |                                                            
     +--+                             +-- Amblema_plicata **            
        |                          +--+                              
        |                          |  +-- Amblema_neislerii **          
        |  +-----------------------+                                 
        |  |                       |  +-- Reginaia_ebenus **            
        |  |                       +--+                              
        |  |                          +-- Reginaia_rotulata **          
        |  |                                                         
        |  |           +----------------- Popenaias_popeii **           
        |  |           |                                             
        |  |  +--------+  +-------------- Psoronaias_guatemalensis[2] **
        |  |  |        |  |                                          
        +--+  |        +--+  +----------- Psoronaias_guatemalensis[1] **
           |  |           |  |                                       
           |  |           +--+  +-------- Psoronaias_semigranosa[3] **  
           |  |              |  |                                    
           |  |              |  |  +----- Psoronaias_semigranosa[2] **  
           |  |              +--+  |                                 
           |  |                 |  +----- Psoronaias_usumasintae[2] **  
           |  |                 |  |                                 
           |  |                 +--+----- Psoronaias_percompressa **    
           +--+                    |                                 
              |                    |  +-- Psoronaias_semigranosa[1] **  
              |                    |  |                              
              |                    +--+-- Psoronaias_profunda **        
              |                       |                              
              |                       +-- Psoronaias_usumasintae[1] **  
              |                                                      
              |                       +-- Cyrtonaias_tampicoensis **    
              |  +--------------------+                              
              |  |                    +-- Glebula_rotundata **          
              |  |                                                   
              +--+  +-------------------- Potamilus_alatus **           
                 |  |                                                
                 |  |                 +-- Truncilla_truncata **         
                 +--+  +--------------+                              
                    |  |              +-- Ptychobranchus_fasciolaris ** 
                    |  |                                             
                    +--+  +-------------- Pachynaias_spheniopsis **     
                       |  |                                          
                       |  |           +-- Cyrtonaias_explicata **       
                       +--+  +--------+                              
                          |  |        +-- Sphenonaias_microdon **       
                          |  |                                       
                          +--+        +-- Lampsilis_cardium **          
                             |  +-----+                              
                             |  |     +-- Ortmanniana_ligamentina **    
                             +--+                                    
                                |  +----- Cambarunio_iris **            
                                +--+                                 
                                   |  +-- Epioblasma_triquetra **       
                                   +--+                              
                                      +-- Ligumia_recta **              

Wu et al., 2022 [Publ 4238], fig. 2d

iT tT bi riS oT roT roS iD
39 39 37 1 0 0 0 80.0
  +-------------------------------- Lepidodesma_languilati **      
  |                                                             
  |                          +----- Lanceolaria_gladiola **        
  |           +--------------+                                  
  |           |              |  +-- Lanceolaria_grayii **          
  |           |              +--+                               
  |           |                 +-- Lanceolaria_lanceolata **      
  |  +--------+                                                 
  |  |        |  +----------------- Acuticosta_chinensis **        
  |  |        |  |                                              
  |  |        |  |        +-------- Cristaria_plicata **           
  |  |        |  |        |                                     
--+  |        +--+  +-----+     +-- Sinanodonta_woodiana **        
  |  |           |  |     |  +--+                               
  |  |           |  |     |  |  +-- Sinanodonta_lucida **          
  |  |           |  |     +--+                                  
  |  |           |  |        |  +-- Anemina_arcaeformis **         
  |  |           +--+        +--+                               
  |  |              |           +-- Anemina_euscaphys **           
  |  |              |                                           
  |  |              |           +-- Anodonta_cygnea **             
  |  |              |  +--------+                               
  |  |              |  |        +-- Anodonta_anatina **            
  |  |              +--+                                        
  +--+                 |  +-------- Platynaias_compressa **        
     |                 +--+                                     
     |                    |  +----- Pyganodon_grandis **           
     |                    +--+                                  
     |                       |  +-- Utterbackia_imbecillis **      
     |                       +--+                               
     |                          +-- Utterbackia_peninsularis **    
     |                                                          
     |                    +-------- Aculamprotula_tortuosa **      
     |                    |                                     
     |  +-----------------+     +-- Aculamprotula_tientsinensis ** 
     |  |                 |  +--+                               
     |  |                 |  |  +-- Aculamprotula_coreana **       
     |  |                 +--+                                  
     |  |                    |  +-- Aculamprotula_polysticta **    
     |  |                    +--+                               
     |  |                       +-- Aculamprotula_scripta **       
     +--+                                                       
        |                 +-------- Unio_tumidus **                
        |  +--------------+                                     
        |  |              |  +----- Unio_crassus **                
        |  |              +--+                                  
        |  |                 |  +--